Belenois

Belenois Hübner, [1819] Caper Whites

Belenois Hübner, [1819]; Verz. bek. Schmett. (6): 92; TS: Papilio calypso Drury
Anaphaeis Hübner, [1819]; Verz. bek. Schmett. (6): 93; TS: Papilio creona Cramer
Glycestha Billberg, 1820; Enum. Ins. Mus. Billb.: 76; TS: Papilio coronea Cramer
Pseudohuphina Stoneham, 1940; Bull. Stoneham Mus. (40): 4; TS: Pieris raffrayi Oberthür
Pseudanaphaeis Bernardi, 1953; Rev. franc. Ent. 20 (1): 50; TS: Pieris gidica Godart

Anaphaeis (Anthocharini); Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 26
Anapheis; [NHM-BMW]
Anapheis[sic, Anaphaeis]; [BAUR], 153
Annaphaeis; Butler, 1870, (missp.); [NHM-BMW]
Belenois; Winhard, 2000, Butterflies of the world 10: 27; [AfrL]; [CoL] ; [iNaturalist] ; [NHM-BMW]
Belenois (Pierinae); Moore, [1881], Lepid. Ceylon 1 (4): 137; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 112
Glycestha; [NHM-BMW]
Pseudanaphaeis; [NHM-BMW]
Pseudohuphina; [NHM-BMW]
?Adelpha[sic, Anaphaeis?] (Pierini); Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 26

The pseudohuphina species-group

Belenois raffrayi (Oberthür, 1878)

Ethiopia, Sudan, W.Kenya, Zaire, Rwanda, W.Tanzania. See [maps] Raffray's White

Pieris raffrayi Oberthür, 1878; Étud. d'Ent. 3: 17, pl. 1, f. 3

Belenois raffrayi; [EBW]; [BK]; [AfrL]; [CoL] ; [iNaturalist]
Pseudohuphina raffrayi;

B. r. raffrayi S.Ethiopia, S.Sudan

Belenois raffrayi raffrayi; [BAFR], 76 (text); [AfrL]; [iNaturalist]

B. r. extendens (Joicey & Talbot, 1927) W.Kenya, Uganda, Rwanda, Burundi, E.Zaire, W.Tanzania?

Belenois raffrayi extendens; [BAFR], 76; [BK], 138; Winhard, 2000, Butterflies of the world 10: 28, pl. 44, f. 12; [AfrL]; [iNaturalist]

B. r. similis Kielland, 1978 W.Tanzania

Belenois raffrayi similis Kielland, 1978; Tijdschr. Ent. 121 (4): 163, pl. 7, f. 25-26; TL: Tanzania, Mpanda, Mt. Sitebi, 2000m

Belenois raffrayi similis; [AfrL]; [iNaturalist]

Belenois margaritacea Sharpe, 1891

Kenya, Tanzania. See [maps] Margarita's Caper White

Belenois margaritacea Sharpe, 1891; Proc. Zool. Soc. Lond. 1891: 191, pl. 16, f. 4; TL: Sotik

Belenois margaritacea; [BK], 138; [AfrL]; [CoL] ; [iNaturalist]
Pseudohuphina margaritacea;

B. m. margaritacea SW.Kenya (Sotik)

Belenois margaritacea margaritacea; [BAFR], 76; [BK], 138 (note); [AfrL]; [iNaturalist]

B. m. kenyensis (Joicey & Talbot, 1927) SE.Kenya (Teita Hills, Chyulu Hills)

Belenois margaritacea kenyensis; [BAFR], 76; [BK], 138 (note); [AfrL]; [iNaturalist]

B. m. plutonica (Joicey & Talbot, 1927) N.Tanzania (Mt. Kilimanjaro, Usambara Mts., Ngorongoro Crater)

Belenois margaritacea plutonica; [BAFR], 76; [AfrL]; [iNaturalist]

B. m. somereni (Talbot, 1928) Kenya (Meru, Matthew's Range)

Pieris margaritacea somereni Talbot, 1928; Bull. Hill. Mus. 2: 29; TL: Meru

Belenois margaritacea somereni; [BK], 138 (note); [AfrL]; [iNaturalist]

B. m. intermedia Kielland, 1982

Belenois margaritacea intermedia; [AfrL]; [iNaturalist]

Belenois albomaculatus (Goeze, 1779)

Papilio albomaculatus Goeze, 1779; Ent. Beyträge 3 (1): 182

Belenois albomaculatus; [AfrL]; [CoL] ; [iNaturalist]

Belenois grandidieri (Mabille, 1878)

Madagascar, Aldabra Is.. See [maps]

Pieris grandidieri Mabille, 1878; ; TL: Madagascar

Belenois (Belenois) grandidieri; [MadL]
Belenois grandidieri; [BAFR], 76; Winhard, 2000, Butterflies of the world 10: 27, pl. 43, f. 6; [AfrL]; [iNaturalist] ; [LEPIFORUM]

Belenois larima (Boisduval, 1836)

Senegal?, Congo. See [maps]

Pieris larima Boisduval, 1836; Hist. nat. Ins., Spec. gén. Lépid. 1: 524

Belenois larima; [BAFR], 82 (text); [AfrL]; [CoL] ; [iNaturalist]

Belenois zochalia (Boisduval, 1836)

Cape - Moçambique, Rhodesia - Ethiopia, Uganda. See [maps] Forest Caper White Forest White

Pieris zochalia Boisduval, 1836; Hist. nat. Ins., Spec. gén. Lépid. 1: 506
Pieris hyoma Boisduval, 1836; Hist. nat. Ins., Spec. gén. Lépid. 1: 508

Belenois zochalia; [PBSA]; [AfrL]; [CoL] ; [iNaturalist] ; [LEPIFORUM]

B. z. zochalia South Africa, Rhodesia

Belenois zochalia zochalia; [BAFR], 76; [AfrL]; [iNaturalist]

B. z. agrippinides (Holland, 1896) Malawi, Tanzania, Kenya, Uganda

Pieris agrippinides Holland, 1896; Proc. U.S. Nat. Mus. 18 (1098): 758

Belenois zochalia agrippinides; [BAFR], 76; [BK], 139; Winhard, 2000, Butterflies of the world 10: 27, pl. 43, f. 12; [AfrL]; [iNaturalist]

B. z. tanganjikae Lanz, 1896

Belenois zochalia var. tanganjikae Lanz, 1896; Dt. ent. Z. Iris 9 (1): 123
Belenois formosa Butler, 1898; Trans. ent. Soc. 1898: 434

B. z. connexiva (Joicey & Talbot, 1927) Cameroun (highlands)

Belenois zochalia connexiva; [BAFR], 76 (text); [AfrL]; [iNaturalist]

B. z. galla (Ungemach, 1932) Ethiopia

Belenois zochalia galla; [BAFR], 76 (text); [AfrL]; [iNaturalist]

?B. z. camerounica Bernardi, 1966² Nigeria - Cameroun (hihlands)

Belenois camerounica Bernardi, 1966;
Belenois zochalia camerounica; [BK], 139 (note)

Belenois ?sp.

Belenois ?sp.; Winhard, 2000, Butterflies of the world 10: 27, pl. 43, f. 13 (Mpanda, Ntakatta, Kigoma, Tanzania)

Belenois crawshayi Butler, 1894

Angola, Zambia - E.Zaire, Uganda, W.Kenya, Tanzania. See [maps] Crawshay's Caper White

Belenois crawshayi Butler, 1894; Proc. zool. Soc. Lond. 1893: 665; TL: Lake Mweru
Pieris continentalis Heyn, 1904; Ent. Zs. 18: 57

Belenois crawshayi; [BAFR], 78; [BK], 139; [AfrL]; [CoL] ; [iNaturalist]

Belenois diminuta Butler, 1894

Tanzania. See [maps]

Belenois diminuta Butler, 1894; Proc. zool. Soc. Lond. 1893: 666, pl. 60, f. 7; TL: Lake Mweru

Belenois diminuta; Winhard, 2000, Butterflies of the world 10: 27, pl. 43, f. 14; [AfrL]; [CoL] ; [iNaturalist]

Belenois victoria Dixey, 1915

Uganda, Burundi, Rwanda, Tanzania, E.Zaire, W.Kenya. See [maps] Victoria White

Belenois victoria Dixey, 1915; Trans. Ent. Soc. 1915 (1): 4, pl. 2, f. 5-8; TL: Tiriki, Victoria Nyanza; Toro, W.Uganda

Belenois victoria; [BK]; [AfrL]; [CoL] ; [iNaturalist]

B. v. victoria W.Kenya (highlands, W of Rift Valley), Uganda

Belenois victoria victoria; [BK], 139; [BAFR], 78; [AfrL]; [iNaturalist]

B. v. hecqi Berger, 1953 NE.Zaire

Belenois victoria hecqi; [BAFR], 78 (text); [AfrL]; [iNaturalist]

B. v. schoutedeni Berger, 1953 W.Uganda, E.Zaire (Kivu)

Belenois victoria schoutedeni; [BAFR], 78; Winhard, 2000, Butterflies of the world 10: 27, pl. 43, f. 15, 18; [AfrL]; [iNaturalist]

Belenois subeida (C. & R. Felder, 1865)

Guinea - Ethiopia, W.Kenya, Tanzania. See [maps]

Belenois instabilis Butler, 1888; Proc. Zool. Soc. Lond. 1888: 76; TL: Fóda
Belenois subeida ab. lutea Rebel, 1914; Ann. Mus. Wien. 28: 227
Belenois subeida ab. unilutea Rebel, 1914; Ann. Mus. Wien. 28: 227
Belenois subeida ab. infumata Rebel, 1914; Ann. Mus. Wien. 28: 227

Belenois subeida subeida; [iNaturalist]
Belenois subeida; [AfrL]; [CoL] ; [iNaturalist]

B. s. frobeniusi (Strand, 1909) W.Africa

Belenois subeida frobeniusi; [BAFR], 80 (text); [AfrL]; [iNaturalist]

B. s. hiemalis Ungemach² Ethiopia

Belenois hiemalis Ungemach;
Belenois subeida hiemalis; [BAFR], 80 (text)

B. s. sylvander Grose-Smith, 1890 W.Kenya, Uganda, Tanzania, N.Zaire, C.A.R.

Belenois sylvander Grose-Smith, 1890; Proc. Zool. Soc. Lond. 1890: 464

Belenois subeida sylvander; [BAFR], 80; [BK], 140; Winhard, 2000, Butterflies of the world 10: 27, pl. 42, f. 9; [AfrL]; [iNaturalist]

B. s. hailo (Ungemach, 1932) S.Ethiopia, SE.Sudan

Belenois calypso hailo; [BAFR], 82 (text)
Belenois subeida hailo; [AfrL]; [iNaturalist]

Belenois calypso (Drury, [1773])

Gambia - Angola, W.Kenya, Tanzania. See [maps] Calypso Caper White

Papilio calypso Drury, [1773]; Illust. Nat. Hist. Exot. Insects 2: index, 30, pl. 17, f. 3-4
Papilio nigronotatus Goeze, 1779; Ent. Beyträge 3 (1): 182
Papilio nigropictus Goeze, 1779; Ent. Beyträge 3 (1): 183

Belenois calypso; [EBW]; [AfrL]; [CoL] ; [iNaturalist] ; [LEPIFORUM]

B. c. calypso Nigeria - Ghana, Ivory Coast, Liberia, Sierra Leone

Belenois calypso calypso; [BAFR], 82; [AfrL]; [iNaturalist]

B. c. dentigera Butler, 1888 Cameroun - Zaire, NW.Angola

Belenois dentigera Butler, 1888; Proc. Zool. Soc. Lond. 1888: 78; TL: Angola, Stazione Gadda

Belenois calypso dentigera; [BAFR], 82 (text); Winhard, 2000, Butterflies of the world 10: 27, pl. 42, f. 5-6; [AfrL]; [iNaturalist]

B. c. minor Talbot, 1943 Uganda, W.Kenya (W of Rift Valley)

Belenois calypso minor; [BAFR], 82 (text); [BK], 140; [AfrL]; [iNaturalist]

B. c. marlieri Berger, 1981

Belenois calypso marlieri; [AfrL]; [iNaturalist]

Belenois welwitschii Rogenhofer, 1890

-. See [maps]

?Pieris dentigera ratidengi Suffert, 1904; Dt. Ent. Z. Iris 17 (1): 128; TL: Lindi, Deutsch-Ost-Africa

Belenois welwitschii; [AfrL]; [iNaturalist]

B. w. welwitschii NE.Angola, S.Zaire (Shaba), Zambia, Malawi, S.Tanzania, W.Tanzania

Belenois calypso welwitschii; [BAFR], 82
Belenois welwitschii welwitschii; [AfrL]; [iNaturalist]

B. w. shaba Berger, 1981

Belenois calypso shaba Berger, 1981

Belenois calypso shaba; Winhard, 2000, Butterflies of the world 10: 27, pl. 42, f. 7-10
Belenois welwitschii shaba; [AfrL]; [iNaturalist]

Belenois sudanensis (Talbot, 1929)

Cameroun - W.Kenya, Tanzania. See [maps] Sudan Caper White

Belenois sudanensis; [BK], 140; [AfrL]; [CoL] ; [iNaturalist]

B. s. sudanensis (Talbot, 1929)

Belenois sudanensis sudanensis; [AfrL]; [iNaturalist]

B. s. pseudodentigera Berger, 1981

Belenois sudanensis pseudodentigera; Winhard, 2000, Butterflies of the world 10: 27, pl. 42, f. 12; [AfrL]; [iNaturalist]

B. s. katalensis Berger, 1981

Belenois sudanensis katalensis; Winhard, 2000, Butterflies of the world 10: 27, pl. 42, f. 15; [AfrL]; [iNaturalist]

B. s. mayumbana Berger, 1981

Belenois sudanensis mayumbana; [AfrL]; [iNaturalist]

Belenois rubrosignata (Weymer, 1901)

Cameroun - S.Sudan, Uganda, W.Kenya, Tanzania. See [maps] Red-Edged White

Belenois rubrosignata; [EBW]; [AfrL]; [CoL] ; [iNaturalist]

B. r. rubrosignata Angola, S.Zaire (Shaba), Zambia

Belenois rubrosignata rubrosignata; [BAFR], 80; [BK], 141; [AfrL]; [iNaturalist]

B. r. kongwana Talbot, 1943 Malawi, Tanzania, S.Uganda

Belenois rubrosignata kongwana; [BAFR], 80 (text); Winhard, 2000, Butterflies of the world 10: 27, pl. 44, f. 2-3; [AfrL]; [iNaturalist]

B. r. peeli Dixey, 1900 Kenya, Somalia

Belenois rubrosignata peeli; [AfrL]; [BAFR], 80 (text); [iNaturalist]

Belenois thysa (Hopffer, 1855)

Natal, Moçambique, Rhodesia - Kenya, Ethiopia, Somalia. See [maps] False Dotted Border

Pieris thysa Hopffer, 1855; Ber. Verh. Akad. Berlin 1855: 639
?Pieris sabrata Doubleday, [1847]; Gen. diurn. Lep. (1): 47; TL: Congo
?Belenois sabrata Butler, 1870; Trans. ent. Soc. Lond. 1870 (4): 526
Pieris larima rimala Suffert, 1904; Dt. Ent. Z. Iris 17 (1): 80; TL: Bugundi, S. Albert Njansa
Pieris thysa malaria Suffert, 1904; Dt. Ent. Z. Iris 17 (1): 81; TL: Loango
Pieris larima alarmi Suffert, 1904; Dt. Ent. Z. Iris 17 (1): 81; TL: Muanza

Belenois thysa; [EBW]; [PBSA]; [BK], 141; [AfrL]; [CoL] ; [iNaturalist] ; [LEPIFORUM]
thysa f. malaria; [wet season f.]

B. t. thysa E.Cape - Natal, Mozambique, Rhodesia, Zambia, Malawi, Tanzania, Kenya (coast)

Belenois thysa thysa; [BAFR], 82; Winhard, 2000, Butterflies of the world 10: 27, pl. 43, f. 16, pl. 44, f. 1, back f. 5; [AfrL]; [iNaturalist]

B. t. tricolor Talbot, 1943 Ethiopia

Belenois thysa tricolor; [BAFR], 82; [AfrL]; [iNaturalist]

B. t. meldolae Butler, 1872 Angola, S.Zaire - Uganda, W.Kenya, S.Sudan

Belenois meldolae Butler, [1872]; Proc. zool. Soc. Lond. 1871 (3): 725

Belenois thysa meldolae; [BAFR], 82 (text); [AfrL]; [iNaturalist]

Belenois solilucis Butler, 1874

S.Ethiopia, S.Sudan - Uganda, NW.Tanzania, W.Kenya, Cameroun, Angola. See [maps]

Belenois solilucis Butler, 1874; Trans. ent. Soc. Lond. 1874 (4): 433; TL: Angola
Phrissura narcissus Butler, 1898; Trans. ent. Soc. 1898: 431; TL: Angola
Pieris glucki Suffert, 1904; Dt. Ent. Z. Iris 17 (1): 77; TL: Mukenge
Pieris abti Suffert, 1904; Dt. Ent. Z. Iris 17 (1): 77; TL: Central Africa
Pieris solilucis f. sabulosa Joicey & Talbot, 1921; Bull. Hill Mus. 1 (1): 45; TL: Cartouche Village, near Lesse, W bank of Semliki

Belenois solilucis; [EBW]; [BK], 141; [AfrL]; [CoL] ; [iNaturalist]

B. s. solilucis N.Angola, Zaire - W.Tanzania, Cameroun

Belenois solilucis solilucis; [BAFR], 85 (text); [AfrL]; [iNaturalist]

B. s. agylla (Rogenhofer, 1889)

solilucis agylla (Rogenhofer, 1889) (Mylothris); Ann. Mus. Wien. 4: 549, pl. 23, f. 4
Pieris sulphurescens Capronnier, 1889; Bull. Ent. Soc. Belg. 33: cxx

B. s. loveni (Aurivillius, 1921) Uganda, NE.Zaire, S.Sudan, SW.Ethiopia

Belenois solilucis loveni; [BAFR], 85; Winhard, 2000, Butterflies of the world 10: 28, pl. 44, f. 10-11; [AfrL]; [iNaturalist]

Belenois aldabrensis (Holland, 1896) Aldabra, Astove, Assumption Is.

Teracolus aldabrensis Holland, 1896; Proc. U.S. Nat. Mus. 18 (1064): 269, pl. 8, f. 7-8

Belenois aldabrensis; [AfrL]; [BAFR], 85; [CoL] ; [iNaturalist]
Teracolus aldabrensis; Butler, 1897, Ann. Mag. nat. Hist. (6) 20 (119): 464

Belenois anomala (Butler, 1881) Socotra I.

Synchloe anomala Butler, 1881; Proc. zool. Soc. Lond. 1881: 178, pl. 18, f. 3; TL: Sokotra

Belenois anomala; [AfrL]; [CoL] ; [iNaturalist]
Belenois anomalis[sic?]; [BAFR], 86

?Belenois ogygia (Trimen, 1883)

Natal. See [maps]

Pieris ogygia Trimen, 1883; Trans. ent. Soc. Lond. 1883 (4): 356

Belenois ogygia; [BAFR], 79; [CoL] ; [iNaturalist]

Belenois antsianaka (Ward, 1870)

Madagascar. See [maps]

Pieris antsianaka Ward, 1870; Ent. mon. Mag. 7: 30; TL: Madagascar
Pieris ramona Grose-Smith, 1891; Ann. Mag. nat. Hist. (6) 7 (37): 123; TL: NW.Madagascar
Nepheronia[sic] antsianaka var. lasti Grose-Smith & Kirby, 1892; Rhop. Exot. [1] 1: (Belenois) pl. 1, f. 4-5 (preocc.,nom. nud.?)
Belenois antsianaka var. drurii Grose-Smith & Kirby, 1892; Rhop. Exot. [1] 1: (Belenois) 1 (repl. Nepheronia antsianaka var. lasti Grose-Smith & Kirby, 1892); TL: Slary, Madagascar
Belenois antsianaka var. persimilis Grose-Smith & Kirby, 1892; Rhop. Exot. [1] 1: (Belenois) 2, pl. 1, f. 6-7; TL: Salary, Madagascar

Belenois (Belenois) antsianaka; [MadL]
Belenois antsiaka[sic?]; [EBW]; [BAFR], 86
Belenois antsianaka; Grose-Smith & Kirby, 1892, Rhop. Exot. [1] 1: (Belenois) 1; Winhard, 2000, Butterflies of the world 10: 28, pl. 44, f. 4; [AfrL]; [CoL] ; [iNaturalist]
Belenois ramona; Grose-Smith & Kirby, 1892, Rhop. Exot. [1] 1: (Belenois) 2, pl. 1, f. 1-3

Belenois mabella Grose-Smith, 1891

Madagascar. See [maps]

Belenois mabella Grose-Smith, 1891; Ann. Mag. nat. Hist. (6) 8 (43): 79; TL: Madagascar
Belenois mahobo Grose-Smith, 1891; Ann. Mag. nat. Hist. (6) 8 (43): 80; TL: Madagascar

Belenois (Pseudanaphaeis) mabella; [MadL]
Belenois mabella; [BAFR], 86; Winhard, 2000, Butterflies of the world 10: 28, pl. 44, f. 9; [AfrL]; [CoL] ; [iNaturalist]
Pinacopteryx mabella; Grose-Smith & Kirby, 1893, Rhop. Exot. [1] 2: (Pinacopteryx) 1, pl. 1, f. 4-5
Pinacopteryx mahobo; Grose-Smith & Kirby, 1893, Rhop. Exot. [1] 2: (Pinacopteryx) 1, pl. 1, f. 1-3

Belenois java (Linnaeus, 1768)

Indonesia - Samoa, Australia. See [maps] Caper White

Papilio java Linnaeus, 1768; Amoenitates Acad. 7: 504 nota
Papilio coronea Cramer, [1775]; Uitl. Kapellen 1 (1-7): 106, pl. 68, f. B, C
Papilio clytie Donovan, 1805; Epitome nat. Hist. Ins. New Holland: pl. 19, f. 2
Papilio deiopea Donovan, 1805; Epitome nat. Hist. Ins. New Holland: pl. 21, f. 2
Pieris niseia MacLeay, [1826]; in King, Narr. Surv. Austr. 2: 459 ([?] niscia)
Belenois clarissa Butler, 1883; Ann. Mag. nat. Hist. (5) 12 (72): 390; TL: Viti Is.
Belenois java ab. magniplaga Fruhstorfer, 1898; Berl. Ent. Zs. 42 (3/4): 325; TL: Sumba
Anaphaeis nigrita Joicey & Talbot, 1928

Anaphaeis java; [BOW]: pl. 156, f. 11
Anapheis java; [EBW]; [BOR], 144; [BAUR]
Belenois java; Dunn & Dunn, 1991, Review Austr. Butts.: (120-140) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 113; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Don Herbison-Evans]
Belenois java -:; Don Herbison-Evans
Belenois niseia clarissa; Fruhstorfer, 1902, Stettin ent. Ztg 63 (1): 356
Pieris java; Piepers & Snellen, 1909, Rhop. Java [1]: 4

B. j. java Java, Kangean - Sumbawa, Sumba, Sulawesi

Anapheis java java; [BOR], 144
Belenois java java; Winhard, 2000, Butterflies of the world 10: 27, pl. 43, f. 10; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 113

?ssp.

Belenois java ?ssp.; Winhard, 2000, Butterflies of the world 10: 27, pl. 43, f. 11 (Gaibobo I., Papua New Guinea)

B. j. savuana (Fruhstorfer, 1898) Sawu I. (Savu)

Pieris (Belenois) teutonia savuana Fruhstorfer, 1898; Berl. Ent. Zs. 42 (3/4): 326; TL: Savu I.

Anapheis java savuana; [BOR], 144

B. j. teutonia (Fabricius, 1775) Australia

Papilio teutonia Fabricius, 1775; Syst. Ent.: 474 [?]199

Anapheis java teutonia; [BAUR], 153
Belenois java teutonia; [iNaturalist]

B. j. ina Fruhstorfer² Timor?, Wetar, Batar, Kissar

Belenois teutonia anita Fruhstorfer, 1903; Berl. ent. Zs. 48 (1/2): 107; TL: Wetter I.

Anapheis java ina; [BAUR], 153 (text)

B. j. peristhene (Boisduval, 1859) New Hebrides, New Caledonia, Loyalty Is., Norfolk I., Woodlark I.

Pieris peristhene Boisduval, 1859; Bull. Soc. ent. Fr. (3) 7: clv

Anaphaeis java peristhene; Holloway & Peters, 1976, J. Nat. Hist. 10: 290
Anapheis java peristhene; [BAUR], 153 (text)
Belenois java peristhene; [iNaturalist]

B. j. picata Butler, 1882 New Britain, New Ireland, Duke of York I.

Belenois picata Butler, 1882; Ann. Mag. Nat. Hist. (5) 10 (56): 153; TL: New Britain

Anapheis java picata; [BAUR], 153 (text)

B. j. micronesia (Fruhstorfer, 1902) Fiji

Pieris teutonia micronesia Fruhstorfer, 1902; Stettin ent. Ztg 63 (1): 357; TL: Viti
Pieris peristhene vitiensis Fruhstorfer, 1902; Stettin ent. Ztg 63 (1): 358; TL: Viti

Anapheis java micronesia; [BAUR], 153 (text)

Belenois helcida (Boisduval, 1833)

Madagascar. See [maps]

Pieris helcida Boisduval, 1833; Faun. ent. Madagascar, Lépid.: 17, pl. 2, f. 1-2; TL: Madagascar

Belenois (Belenois) helcida; [MadL]
Belenois helcida; [BAFR], 84; [AfrL]; [CoL] ; [iNaturalist] ; [LEPIFORUM]

Belenois hedyle (Cramer, [1777])

-. See [maps]

Papilio hedyle Cramer, [1777]; Uitl. Kapellen 2 (9-16): 137, pl. 186, f. C, D
?Papilio sordide-flavicans Goeze, 1779; Ent. Beyträge 3 (1): 182

Belenois hedyle; [AfrL]; [CoL] ; [iNaturalist] ; [LEPIFORUM]

B. h. hedyle Sierra Leone, Liberia, Ivory Coast, Ghana, Benin

Belenois hedyle hedyle; [BAFR], 84 (text); [AfrL]; [iNaturalist]

B. h. rhena (Doubleday, 1846) Ghana - W.Nigeria

Pieris rhena Doubleday, 1846; Ann. Mag. nat. Hist. 17 (109): 24

Belenois hedyle rhena; [BAFR], 84; [AfrL]; [iNaturalist]

B. h. ianthe (Doubleday, 1842) Sierra Leone, Togo, Guinea, Guinea-Bissau

Pieris ianthe Doubleday, 1842; in Gray, Zool. Misc.: 77

Belenois hedyle ianthe; [AfrL]; [iNaturalist]
Belenois ianthe; [BAFR], 84; [CoL]

Belenois theora (Doubleday, 1846)

-. See [maps]

Pieris theora Doubleday, 1846; Ann. Mag. nat. Hist. 17 (109): 25
Pieris paroreia Hewitson, 1869; Trans. ent. Soc. Lond. 1869 (1): 72; TL: Cape Coast
Pieris lortzingi Suffert, 1904; Dt. Ent. Z. Iris 17 (1): 79; TL: Sesse I., Deutch-Ost-Afrika

Belenois theora; [EBW]; [AfrL]; [CoL] ; [iNaturalist] ; [LEPIFORUM]

B. t. theora Sierra Leone - Nigeria

Belenois theora theora; [BAFR], 81; [AfrL]; [iNaturalist]

?B. t. concolor Aurivillius² Cameroun - C.A.R., C.Zaire, S.Zaire

Belenois concolor Aurivillius;
Belenois theora concolor; [BAFR], 81 (text)

B. t. ratheo (Suffert, 1904) C.A.R

Pieris theora ratheo Suffert, 1904; Dt. Ent. Z. Iris 17 (1): 81; TL: Johann Albrechthöhe, Camerun

Belenois theora ratheo; Winhard, 2000, Butterflies of the world 10: 28, pl. 44, f. 7-8; [AfrL]; [iNaturalist]

B. t. laeta (Weymer, 1903) W.Uganda, E.Zaire, S.Sudan

Belenois theora laeta; [BAFR], 81 (text); [AfrL]; [iNaturalist]

Belenois theuszi (Dewitz, 1889)

S.Angola, NW.Zambia, S.Zaire - Gabon, Cameroun, W.Nigeria. See [maps]

?Pieris theuszi var. semialba Aurivillius, 1895; Ent. Tidskr. 16: 261

Belenois theuszi; [BAFR], 82; Winhard, 2000, Butterflies of the world 10: 28, pl. 43, f. 17; [AfrL]; [CoL] ; [iNaturalist]

Subgenus Anaphaeis Hübner, [1819]

Anaphaeis Hübner, [1819]; Verz. bek. Schmett. (6): 93

Anaphaeis; [NHM-BMW]

Belenois aurota (Fabricius, 1793)

Ceylon, India, NW.India, Pakistan, Afghanistan, Baluchistan, Arabia, SAF, Tropical Africa (dry zones). See [maps] Brown-veined White Caper White Pioneer

Papilio aurota Fabricius, 1793; Ent. Syst. 3 (1): 197; TL: Tranquebar, S.India
Papilio mesentina Cramer, [1780]; Uitl. Kapellen 3 (23-24): 140, pl. 270, f. A, B (preocc. Papilio mesentina Cramer, [1777])
Pinacopteryx syrinx Wallengren, 1863; Wien ent. Monats. 4 (2): 34

Anaphaeis aurota; [BMAT]: 18, pl. 6, f. 27-29; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 26
Anapheis aurota; [BRU], 160
Anapheis[sic] mesentina; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 26
Belenois aurota; [EBW]; [PBSA]; [BOR], 146; [AfrL]; [CoL] ; [iNaturalist] ; [LEPIFORUM]
mesentina; [BIR], 425

745x923 (~326Kb) upperside Ethiopia, Amhara, Jara River shingle bank at the bridge 9 km SSW of Hayk. 6th August 2012, Photo © Oleg Kosterin

972x935 (~394Kb) underside Ethiopia, Amhara, Jara River shingle bank at the bridge 9 km SSW of Hayk. 6th August 2012, Photo © Oleg Kosterin

B. a. aurota India, Sikkim, Tropical Africa, Madagascar

Belenois (Belenois) aurota aurota; [MadL]
Belenois aurota aurota; [BOR], 146; [BAFR], 78; [BK], 137; Winhard, 2000, Butterflies of the world 10: 27, pl. 42, f. 13, pl. 43, f. 1; [iNaturalist]
?mesentina r. mesentina; [BIR], 425

B. a. augusta (Olivier, 1801)

Papilio augusta Olivier, 1801; Voy. L'Emp. Othoman. Atl. 4 (1801?): pl. 33, f. 3a-b

Belenois aurota augusta; Winhard, 2000, Butterflies of the world 10: 27, pl. 43, f. 4

B. a. taprobana (Moore, 1872) Ceylon

Pieris taprobana Moore, 1872; Proc. zool. Soc. Lond. 1872 (2): 565; TL: Ceylon
Belenois mesentina fervidior Fruhstorfer, 1898; Berl. Ent. Zs. 42 (3/4): 326; TL: Ceylon

Belenois aurota taprobana; [BOR], 146; [iNaturalist]
Belenois taprobana; Moore, [1881], Lepid. Ceylon 1 (4): 137, (3): pl. 53, f. 3, 3a
mesentina r. taprobana; [BIR], 425

?ssp.

Belenois aurota ?ssp.; Winhard, 2000, Butterflies of the world 10: 27, pl. 42, f. 14, pl. 43, f. 2-3 (Lmandraka, Madagascar)

B. a. turanica (Sheljuzhko, 1908) S.Turkmenia, Tajikistan

aurota turanica (Sheljuzhko, 1908) (Pontia); Rev. russ. ent. 7 (4): 233

Adelpha[sic] mesentina turanica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 26

Belenois creona (Cramer, [1776])

Tropical Africa (dry parts), Comoros, Madagascar, SW.Arabia. See [maps] African Common White African Caper

Papilio creona Cramer, [1776]; Uitl. Kapellen 1 (8): 148, pl. 95, f. B, C
Colias ernestius Laporte, 1830; Actes Soc. Linn. Bordeaux 4: 147

Belenois creona; [EBW]; [AfrL]; [CoL] ; [iNaturalist] ; [LEPIFORUM]

B. c. creona Senegal - Nigeria, Sudan, W.Ethiopia

Belenois creona creona; [BAFR], 79 (text); [AfrL]; [iNaturalist]

B. c. benadirensis (Storace, 1948) Somalia

Belenois creona benadirensis; [BAFR], 79 (text); [AfrL]; [iNaturalist]

B. c. boguensis (C. & R. Felder, 1865) Agamsa?, Ethiopia

Pieris boguensis C. & R. Felder, 1865; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 173

Belenois creona boguensis; [BAFR], 79 (text); [AfrL]; [iNaturalist]

B. c. elisa (Vollenhoven, 1869) Comoro Is.

Pieris elisa Vollenhoven, 1869; Faune Madag. 5 (Ins.): 12

Belenois creona elisa; [BAFR], 79 (text); [AfrL]; [iNaturalist]

B. c. leucogyne Butler, 1885 SW.Arabia

Belenois creona leucogyne; [BAFR], 79 (text); [AfrL]; [iNaturalist]

B. c. prorsus (Talbot, 1943) Madagascar

Anapheis creona prorsus Talbot, 1943; Ann. Mag. nat. Hist. (11) 10 (65): 305; TL: Madagascar, Maintirano

Belenois (Belenois) creona prorsus; [MadL]
Belenois creona prorsa; Winhard, 2000, Butterflies of the world 10: 27, pl. 43, f. 5
Belenois creona prorsus; [BAFR], 79 (text); [AfrL]; [iNaturalist]

B. c. severina (Stoll, [1781]) South Africa - E.Africa, Zaire

Papilio severina Stoll, [1781]; in Cramer, Uitl. Kapellen 4 (29-31): 95, pl. 338, f. G, H

Belenois creona severina; [PBSA]; [BAFR], 79; [BK], 137; Winhard, 2000, Butterflies of the world 10: 27, pl. 43, f. 7-9; [AfrL]; [iNaturalist]
Belenois severina; [EBW]

?B. c. infida Butler, 1888

Belenois infida Butler, 1888; Proc. Zool. Soc. Lond. 1888: 78; TL: Wadelai

Belenois gidica (Godart, 1819)

Natal, Swaziland, Transvaal, Moçambique, Rhodesia, South West Africa, Tropical Africa (dry zones). See [maps] Pointed Caper African Veined White

Pieris gidica Godart, 1819; Encyclopédie Méthodique 9 (1): 131
Belenois occidentis Butler, 1898; Trans. ent. Soc. 1898: 438; TL: Congo; Loanda
?Pieris gidica var. westwoodi Wallengren
?Pieris gidica ab. simplex Rebel, 1914; Ann. Mus. Wien. 28: 226

Belenois gidica; [EBW]; [PBSA]; [BAFR], 86; [BK], 138; [AfrL]; [CoL] ; [iNaturalist] ; [LEPIFORUM]

592x608 (~190Kb) Ethiopia, Amhara, a rapidous brook, Blue Nile right tributary just upstream the bridge of the road Addis Ababa - Bahir Dar, 1100 m a.s.l. 1st August 2012, Photo © Oleg Kosterin	666x678 (~189Kb) Ethiopia, Amhara, a rapidous brook, Blue Nile right tributary just upstream the bridge of the road Addis Ababa - Bahir Dar, 1100 m a.s.l. 1st August 2012, Photo © Oleg Kosterin	689x646 (~219Kb) upperside Ethiopia, Amhara, a rapidous brook, Blue Nile right tributary just upstream the bridge of the road Addis Ababa - Bahir Dar, 1100 m a.s.l. 1st August 2012, Photo © Oleg Kosterin
identification uncertain 714x799 (~282Kb) Ethiopia, Amhara, a rapidous brook, Blue Nile right tributary just upstream the bridge of the road Addis Ababa - Bahir Dar, 1100 m a.s.l. 1st August 2012, Photo © Oleg Kosterin	identification uncertain 935x859 (~316Kb) Ethiopia, Amhara, a rapidous brook, Blue Nile right tributary just upstream the bridge of the road Addis Ababa - Bahir Dar, 1100 m a.s.l. 1st August 2012, Photo © Oleg Kosterin

B. g. gidica

Belenois gidica gidica; [AfrL]; [iNaturalist]

B. g. hypoxantha (Ungemach, 1932)

Belenois gidica hypoxantha; [AfrL]; [iNaturalist]
Belenois hypoxantha; [iNaturalist]

B. g. abyssinica (Lucas, 1852) Ethiopia

Pieris abyssinica Lucas, 1852; Revue Mag. Zool. (2) 4 (7): 328

Belenois gidica abyssinica; [AfrL]; [iNaturalist]

Belenois johannae Butler, 1879; Ann. Mag. nat. Hist. (5) 3 (15): 191; TL: Johanna I.

Pieris voeltzkowi Karsch, 1900; Ent. Nachr. 26 (24): 369; TL: W.Madagascar

References:

[AfrL] Mark C. Williams
Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[BHL] The
Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[CoL] Catalogue of Life
Bánki, O., Roskov, Y., Döring, M., Ower, G., Hernández Robles, D. R., Plata Corredor, C. A., Stjernegaard Jeppesen, T., Örn, A., Vandepitte, L., Hobern, D., Schalk, P., DeWalt, R. E., Ma, K., Miller, J., Orrell, T., Aalbu, R., Abbott, J., Adlard, R., Aedo, C., et al. (2024). Catalogue of Life Checklist (Version 2024-03-26); https://doi.org/10.48580/dfz8d
[Don Herbison-Evans] and Stella A. Crossley
Australian Caterpillars and their Butterflies and Moths
[EVERSMANNIA]
Eversmannia contents
[HBN] Francis Hemming, 1937
HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[LAMAS-2005] Gerardo Lamas
Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LAMAS-NEO] Gerardo Lamas
Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[LEPIFORUM]
Lepiforum; Bestimmung von Schmetterlingen (Lepidoptera) und ihren Präimaginalstadien
[NHM-BMW] Natural History Museum
Butterflies and Moths of the World; Generic Names and their Type-species
[Oleg Kosterin] Institue of Cytology & Genetics
Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[iNaturalist]
iNaturalist
[maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[²] This may require parentheses or not (original genus not verified)

Some related literature:

Aurivillius, 1895
Beiträge zur Kenntniss der Insektenfaun von Kamerun. 2. Tagfalter Ent. Tidskr. 16 : 195-220, : 255-268
Bernardi, 1953
Note sur la taxonomie des genres Belenois Hb. et Anaphaeis Hb., [Lep. Pieridae] Rev. franc. Ent. 20 (1) : 47-53
Billberg, 1820
Enumeratio Insectorum Enum. Ins. Mus. Billb. : 1-138
Boisduval, 1833
Faune Entomologique de Madagascar, Bourbon et Maurice. Lépidoptéres Faun. ent. Madagascar, Lépid. : 1-122, pl. 1-16
Boisduval, 1859
[communications] Bull. Soc. ent. Fr. (3) 7 : cliv-clviii
Boisduval, 1836
Histoire Naturelle des Insectes. Species Général des Lépidoptéres. Tome Premier Hist. nat. Ins., Spec. gén. Lépid. 1 : 1-690, pl. 1-24
Butler, [1872]
On a small Collection of Butterflies from Angola Proc. zool. Soc. Lond. 1871 (3) : 721-725
Butler, 1874
Descriptions of some new species and a new genus of diurnal Lepidoptera in the collection of Herbert Druce Esq. Trans. ent. Soc. Lond. 1874 (4) : 423-436, pl. 6
Butler, 1879
On a Collection of Lepidoptera from the Island of Johanna Ann. Mag. nat. Hist. (5) 3 (15) : 186-192
Butler, 1881
On the Lepidoptera collected in Socotra by I. Bayley Balfour Proc. zool. Soc. Lond. 1881 : 175-180, pl. 18
Butler, 1882
Descriptions of new species of Lepidoptera, chiefly from Duke of York Island and New Britain Ann. Mag. Nat. Hist. (5) 10 (55): 36-43, (56): 149-160, (57): 226-238
Butler, 1883
New Lepidoptera from the Viti Islands Ann. Mag. nat. Hist. (5) 12 (72) : 389-391
Butler, 1888
On the Lepidoptera received from Dr. Emin Pascha [from Equatorial Africa] Proc. Zool. Soc. Lond. 1888 : 56-85
Butler, 1894
On two collection of Lepidoptera sent by H. H. Johnston, Esq., C.B., from British Central Africa Proc. zool. Soc. Lond. 1893 : 643-684, pl. 60 (1894)
Butler, 1897
A revision of the Species of Butterflies belonging to the Genus Teracolus Swains. Ann. Mag. nat. Hist. (6) 20 (118): 385-399, (119): 451-473, (120): 495-507
Butler, 1898
On some new species of African Pierinae in the collection of the British Museum, with notes on seasonal forms of Belenois Trans. ent. Soc. 1898 : 431-438
Capronnier, 1889
Liste des Lépidoptères capturés au Congo par Messieurs Thys, Legat, Martini et Machado en 1887 Bull. Ent. Soc. Belg. 33 : cxviii-cxxvii
Cramer, [1775]; Cramer, [1776]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 1 (1-7): 1-132, pl. 1-84 (1775), (8): 133-155, pl. 85-96 (1776)
Cramer, [1777]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 2 (9-16): 1-152, pl. 97-192 (1777)
Cramer, [1780]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 3 (17-21): 1-104, pl. 193-252 (1779), (22): 105-128, pl. 253-264 ([1780]), (23-24): 129-176, pl. 265-288 (1780)
[BAUR]; D'Abrera, 1977
Butterflies of the Australian Region (2nd edn) Butts. Aust. Region: 1-416
[BAFR]; D'Abrera, 1980
Butterflies of the Afrotropical Region: -
[BOR]; D'Abrera, 1982-1986
Butterflies of the Oriental Region I-III
[PBSA]; Dickson & Kroon, 1978
Pennington's Butterflies of Southern Africa
Dixey, 1915
New species and subspecies of Pierinae Trans. Ent. Soc. 1915 (1) : 1-15, pl. 1-2
Donovan, 1805
An Epitome of the Natural History of the Insects of New Holland, New Zealand, New Guinea, Otaheite and other Islands in the Indian, Sothern and Pacific Oceans ... Epitome nat. Hist. Ins. New Holland : pl. [1-41]
Doubleday, 1842
Characters of undescribed Lepidoptera in Gray, Zool. Misc.: 73-78
Doubleday, 1846
Descriptions of new or imperfectly described Diurnal Lepidoptera Ann. Mag. nat. Hist. 17 (109) : 22-26
Doubleday, [1847]
The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846), (1): 19-132, pl. 4*, 18*, 5-25, 28 (3-14) (1847), (1): 133-200, pl. 1*, 26, 27, 29 (15-23) (1848), (1): pl. 31-44 (15-23) (1848), (1): 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849), (1): 243-250 [?], (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850), (2): 327-466, pl. 59, 67-77 (39-50) (1851), (2): 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)
Drury, [1773]
Illustrations of natural history; wherein are exhibited ... Illust. Nat. Hist. Exot. Insects 2 : pl. 1-50, : 1-100 (1837)
Dunn & Dunn, 1991
Review of Australian Butterflies: distribution, life history and taxonomy Review Austr. Butts. (1): 1-196, (2): 197-335, (3): 336-512, (4): 513-660
Butler in Dunning, 1870
Notes on a Collection of Insects sent by Mr. Ansell from South-West Africa Trans. ent. Soc. Lond. 1870 (4) : 521-532
Fabricius, 1775
Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibus, Observationibus Syst. Ent. : 1-832
Fabricius, 1793
Entomologia Systematica emendata et aucta. Ent. Syst. 3 (1) : 1-487
C. & R. Felder, 1865
Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Behilfen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Band 2. Abtheilung 2. Lepidoptera. Rhopalocera Reise Fregatte Novara, Bd 2 (Abth. 2) (1): 1-136, pl. 1-21 (Rhop.) (C. & R. Felder, [1865]), (2): 137-378, pl. 22-47 (Rhop.) (C. & R. Felder, 1865), (3): 379-536, pl. 48-74 (Rhop.) (C. & R. Felder, 1867), (4): 537-548, pl. 75-120, (Inhalts-Verz.) 1-9 (pl. 1-74), (Felder & Rogenhofer, 1874) , (5): pl. 121-140, (erklärung) 1-10 (pl. 75-107), (Inhalts-Verz.) 1-20 (pl. 108-140), (Felder & Rogenhofer, 1875)
Fruhstorfer, 1898
Neue Rhopaloceren aus dem Indo-malayischen Archipel Berl. Ent. Zs. 42 (3/4) : 311-343 (mid July)
Fruhstorfer, 1902
Beitrag zur Kenntniss der Lepidopteren der Viti-Inseln Stettin ent. Ztg 63 (1) : 350-359 (1 July)
Fruhstorfer, 1903
Neue Pieriden und Uebersicht verwandter Formen Berl. ent. Zs. 48 (1/2) : 97-111 (early May)
Godart, 1819
Encyclopédie Méthodique. Histoire naturelle Entomologie, ou histoire naturelle des crustacés, des arachnides et des insectes Encyclopédie Méthodique 9 (1): 3-328 (1819), (2): 329-828 ([1824])
Goeze, 1779
Entomologische Beyträge zu des Ritter Linné zwölften Ausgabe des Natursystems Ent. Beyträge 3 (1) : 1-390
Grose-Smith, 1890
A list of the Butterflies collected by Mr. William Bonny on the Journey with Mr. Stanley from Yambuya on the Aruwimi River through the Great Forest of Central Africa; with Descriptions of nine new Species Proc. Zool. Soc. Lond. 1890 : 463-473
Grose-Smith, 1891
Descriptions of Ten new Species of Butterflies from the North-west Coast of Madagascar, captured by Mr. J. T. Last, in the Collection of Mr. H. Grose Smith Ann. Mag. nat. Hist. (6) 7 (37) : 122-128
Grose-Smith, 1891
Descriptions of Four new Species of Butterflies from South-west Madagascar, captured by Mr. J. T. Last, in the Collection of H. Grose Smith Ann. Mag. nat. Hist. (6) 8 (43) : 78-81
Grose-Smith & Kirby, 1892; Grose-Smith & Kirby, 1893
Rhopalocera Exotica, being Illustrations of New, Rare, and Unfigured Species of Butterflies Rhop. Exot. [1] 1: (Ornithoptera) 1-2, pl. 1 (1887), [1] 2: (Ornithoptera) 3-4, pl. 2 (1892), [1] 3: (Ornithoptera) [5-6] 1-2, pl. 3 (1900), [1] 3: (Ornithoptera) 7-8, pl. 4 (1901), [1] 1: (Papilio) 1-2, pl. 1 (1887), [1] 1: (Papilio) 3-5,7-14, pl. 2-6 (1888), [1] 1: (Papilio) 15-20, pl. 7-9 (1890), [1] 1: (Papilio) 21-25, pl. 10-11 (1891), [1] 2: (Papilio) 27-31, pl. 12-13 (1893), [1] 2: (Papilio) 33-35, pl. 14 (1894), [1] 2: (Papilio) 37-38, pl. 15 (1895), [1] 2: (Papilio) 39-40, pl. 16 (1897), [1] 3: (Papilio) 41-46, pl. 17-19 (1899), [1] 3: (Papilio) 47-48, pl. 20 (1900), [1] 3: (Papilio) 49-50, pl. 21 (1901), [1] 3: (Papilio) 51-55, pl. 22-23 (1902), [1] 1: (Delias) 1-3, pl. 1 (1889), [1] 2: (Delias) 5-7, 9-11, pl. 2-3 (1893), [1] 2: (Delias) 12-18, pl. 4-5 (1895), [1] 2: (Delias) 20-22, pl. 6 (1896), [1] 3: (Delias) 24-26, pl. 7 (1897), [1] 3: (Delias) 28-35, pl. 8-9 (1901), [1] 3: (Delias) 36-38, pl. 10 (1902), [1] 1: (Pieridae) 1-3, pl. 1 (1888), [1] 2: (Pinacopteryx) 1-4, pl. 1 (1893), [1] 1: (Callosune) 1-3, pl. 1 (1889), [1] 3: (Huphina) 1-7, pl. 1-2 (1898), [1] 3: (Huphina) [8],9-10, pl. 3 (1899), [1] 1: (Elodina) 1-4, pl. 1 (1890), [1] 1: (Belenois) 1-4, pl. 1 (1892), [1] 2: (Belenois) 5-8, pl. 2 (1892), [1] 3: (Mylothris) 1-3, 5-7, pl. 1-2 (1900), [1] 1: (Ixias) 1-3, pl. 1 (1888), [1] 2: (Appias) 1-3, pl. 1 (1894), [1] 1: (Pierinae) 5-7, pl. 2 (1889), [1] 2: (Dismorphia) 1-4, pl. 1 (1896), [1] 2: (Dismorphia) 5-9, 11-15, pl. 2-3 (1897), [2] 1: (Asthipa) 1-3, pl. 1 (1890), [2] 2: (Asthipa) 5-6, pl. 2 (1896), [2] 2: (Ravadeba) 7, pl. 1 (1896), [2] 2: (Mycalesis) 1-4, pl. 1 (1895), [2] 2: (Mycalesis) 5-12, pl. 2-3 (1896), [2] 3: (Mycalesis) 13-18, pl. 4-5 (1902), [2] 3: (Libythea) 1-2, pl. 1 (1902), : (Ragadia-Argyronympha-Hypocysta) : 1-6, pl. [1] (1895), [2] 2: (Pedaliodes) 1-5, pl. 1 (1893), [2] 2: (Pedaliodes) 7-9, pl. 2 (1894), [2] 2: (Pedaliodes) 10-14, pl. 3 (1895), [2] 3: (Pedaliodes) 15-20, pl. 4-5, (Oxeoschistus) 1 (1900), [2] 3: (Pronophila) 1-2, pl. 1, (Oxeoschistus) 2-3, pl. 1 (1900), [2] 2: (Elymnias) 1-4, pl. 1 (1896), [2] 3: (Elymnias) 5-6, pl. 2, (Bruasa) 6-7, pl. 1 (1897), [2] 1: (Morpho) 1-2, pl. 1 (1890), [2] 3: (Morphotenaris-Hyantis) 1-3, pl. [1] (1898), [2] 2: (Tenaris) 1-3, 5-7, pl. 1-2 (1894), [2] 2: (Tenaris) 9-11, pl. 3 (1895), [2] 2: (Tenaris) 13-15, pl. 4 (1896), [2] 3: (Tenaris) 17-18, pl. 5 (1898), [2] 1: (Heliconius) 1-3, pl. 1 (1892), [2] 1: (Acraea) 1-6, pl. 1-2 (1889), [2] 1: (Acraea) 7-10, pl. 3 (1892), [2] 2: (Acraea) 11-14, pl. 4 (1892), [2] 2: (Acraea) 15-18, pl. 5 (1894), [2] 3: (Acraea) 19-29, pl. 6-8, (Planema) 29-30, pl. [1] (1901), [2] 1: (Cethosia) 1-2, pl. 1 (1889), [2] 3: (Cirrhochroa) 1, pl. [1], (Cupha) 2-4, pl. [1] (1898), [2] 2: (Terinos) 1-2, pl. 1 (1894), [2] 1: (Thaleropis) 1-3, pl. 1 (1891), [2] 2: (Doleschallia) 1-3,5-7, pl. 1-2 (1893), [2] 2: (Kallima) 1-3, pl. 1 (1892), [2] 1: (Hypolimnas) 1-2, pl. 1 (1891), [2] 3: (Hypolimnas) 3-4, pl. 2 (1897), [2] 3: (Hypolimnas) 5-6, pl. 3 (1898), [2] 1: (Euxanthe) 1-3, pl. 1 (1890), [2] 1: (Hestina) 1-3, pl. 1 (1891), [2] 1: (Cymothoe) 1-4, pl. 1 (1890), [2] 3: (Limenitis) 1-3, pl. 1 (1899), [2] 2: (Neptis) 1-4, pl. 1 (1895), [2] 3: (Neptis) 5-11, pl. 2-3 (1899), [2] 3: (Neptis) 13-17, pl. 4-5 (1900), [2] 1: (Euthalia-Tanaecia) 1-3, pl. 1 (1889), [2] 1: (Euthalia) 5-10, pl. 2-3 (1891), [2] 2: (Euthalia) 11-14, pl. 4 (1894), [2] 3: (Euthalia) 15, pl. 5, (Dichorragia) 16, pl. 1 (1898), [2] 1: (Apatura) 1-3, pl. 1 (1892), [2] 1: (Euphaedra) 1-2, pl. 1 (1889), [2] 1: (Euryphene) 1-3, pl. 1 (1889), [2] 1: (Charaxes) 1-2,[3-4] pl. 1-2 (1887), [2] 1: (Charaxes) [5], pl. 3 (1888), [2] 1: (Charaxes) 7-9, pl. 4 (1890), [2] 1: (Charaxes) 11-13, pl. 5 (1891), [2] 1: (Mynes) 1-2, pl. 1 (1888), [2] 2: (Mynes) 3-6, pl. 2 (1894), [2] 2: (Prothoe) 1-2, pl. 1 (1896), [3] 3: (Erycinidae) 1-10, pl. 1-2 (1902), [3] 2: (Dicallaneura-etc) 1-6, pl. 1 (1897), [3] 3: (Dicallaneura-etc) 7-11, pl. 2 (1901), [3] 1: (Lycaenidae) 1-4,[5-9] 1-5, pl. 1-2 (1887), [3] 1: (Lycaenidae) 11-17, 19-22, pl. 3-5 (1888), [3] 1: (Lycaenidae) 23-25, 27-29, 31-33, pl. 6-8 (1889), [3] 1: (Lycaenidae) 35-48, pl. 9-11 (1890), [3] 1: (Lycaenidae) 49-51,53-64, pl. 12-15 (1891), [3] 2: (Lycaenidae) 65-68, pl. 16 (1892), [3] 1: (Lycaenidae) [69-72] 57-60, pl. 17 (1892), [3] 2: (Lycaenidae) [73-77] 61-65,79-84, pl. 18-19 (1892), [3] 2: (Lycaenidae) 85-107, pl. 20-23 (1893), [3] 2: (Lycaenidae) 109-120, pl. 24-25 (1894), [3] 3: (Lycaenidae) 121-125, pl. 26 (1900), [3] 3: (Lycaenidae) 127-131, 133-141, pl. 27-29 (1901), [3] 3: (Lycaenidae) 143-146, pl. 30 (1902), [3] 2: (Hypochrysops) 1-6, pl. 1 (1894), [3] 2: (Hypochrysops) 7-13, 15-22, pl. 2-3 (1895), [3] 2: (Thysonotis) 23-25, 27-31, pl. 1-2 (1895), [3] 2: (Waigeum) 34-38, pl. 1 (1896), [3] 2: (Thysonotis) 39-46, pl. 3-4 (1896), [3] 2: (Holochila) 1-4, pl. 1 (1896), [3] 2: (Holochila) 5-8, pl. 2 (1897), [3] 2: (Pseudonotis) 1-3, pl. [1], (Epimastidia) 4-5, pl. [1], (Lampides) 5-8, pl. [1] (1897), [3] 2: (Lampides) 9-10, pl. 2, (Thysonotis) 47, pl. 5 (1897), [3] 3: (Holochila) 9-11, pl. 3 (1898), [3] 3: (Arhopala) 1-7, pl. 1-2 (1898), [3] 3: (Arhopala) 9-12, pl. 3, (Ilerda) 13, pl. [1], (Horaga) 14-15, pl. [1] (1902), [3] 3: (Thysonotis) 47x,48-50, pl. 6 (1898), [3] 3: (Hypochrysops) 23-26, pl. 4 (1898), [3] 3: (Holochila) 13-15, pl. 4 (1899), [3] 3: (Hypochrysops) 27-29, pl. 5, (Waigeum) [30], pl. 2 (1899), [3] 3: (Holochila) 17-19, pl. 5 (1899), [3] 3: (Thysonotis) 51-54, pl. 7 (1899), [3] 3: (Jamides) 11-14, pl. 1 (1900), [3] 3: (Hypolycaena-etc) 1-4, pl. [1] (1900), [3] 3: (Hypochrysops) 31-33, pl. 6, (Thysonotis) 55-56, pl. 8 (1900), [3]: (Additions-Corrections) 1-2,[3],vii-x
Heyn, 1904
Pieris continentalis n. sp. Ent. Zs. 18 : 57-58
Hewitson, 1869
Descriptions of new Species of Diurnal Lepidoptera Trans. ent. Soc. Lond. 1869 (1) : 71-75
Holland, 1896
List of the Lepidoptera from Aldabra, Seychelles, and other East African Islands, Collected by Dr. W. L. Abbot Proc. U.S. Nat. Mus. 18 (1064) : 265-273
Holland, 1896
List of the Lepidoptera Collected in East Africa, 1894, by Mr. William Astor Chanler and Lieut. Ludwig von Höhnel Proc. U.S. Nat. Mus. 18 (1098) : 741-767
Holloway & Peters, 1976
The butterflies of New Caledonia and the Loyalty Islands J. Nat. Hist. 10 : 273-318
Hopffer, 1855
[Lepidoptera Diurna] Ber. Verh. Akad. Berlin 1855 : 639-643
Hübner, [1819]
Verzeichniss bekannter Schmettlinge, 1816-[1826] Verz. bek. Schmett. (1): [1-3], 4-16 (1816), (2): 17-32 (1819), (3): 33-48 (1819), (4): 49-64 (1819), (5): 65-80 (1819), (6): 81-96 (1819), (7): 97-112 (1819), (8): 113-128 (1819), (9): 129-144 (1819), (10): 145-160 (1819), (11): 161-176 (1819), (12): 177-192 (1820), (13): 193-208 (1820), (14): 209-224 (1821), (15): 225-240 (1821), (16): 241-256 (1821), (17): 257-272 (1823), (18): 273-288 (1823), (19): 289-304 (1823), (20): 305-320 (1825), (21): 321-336 (1825), (22): 337-352 (1825), (23-27): 353-431 ([1825])
Joicey & Talbot, 1921
New Lepidoptera Collected by Mr. T. A. Barns, in East Central Africa. New Forms of Rhopalocera Bull. Hill Mus. 1 (1) : 44-118, pl. 5-16
Karsch, 1900
Vorläufige kurze Kennzeichnung von fünf neuen, durch Herrn Dr. A. Voelzkow in West-Madagascar entedecten Lepidopteren Ent. Nachr. 26 (24) : 369-370
Kielland, 1978
A provisional checklist of the Rhopalocera of the eastern side of Lake Tanganyika Tijdschr. Ent. 121 (4) : 147-237
Korb & Bolshakov, 2011
[A catalogue of butterflies (Lepidoptera: Papilioformes) of the former USSR. Second edition, reformatted and updated] (in Russian) Eversmannia Suppl. 2 : 1-123
Lanz, 1896
Besprechung der von Dr. Bumiller 1893 aus Ostafrika mitgebrachten Schmetterlinge Dt. ent. Z. Iris 9 (1) : 113-147
Laporte, 1830
Observations pour servir a l'histoire de quelques insectes, et description d'une nouvelle espece de Coliade Actes Soc. Linn. Bordeaux 4 : 141-156
[BK]; Larsen, 1996
The Butterflies of Kenya and their natural history Butterflies of Kenya
[MadL]; Lees, Kremen & Raharitsimba, 2003
Classification, Diversity and Endemism of the Butterflies (Papilionoidea and Hesperioidea): A Revised Species Checklist in Goodman & Benstead, Nat. Hist. Madagascar : 762-793
[BOW]; Lewis, 1974
Butterflies of the World
Lucas, 1852
Description de nouvelles Espèces de Lépidoptères appartenant aux Collections entomologiques du Musée de Paris Revue Mag. Zool. (2) 4 (3): 128-141 (1852), 4 (4): 189-198 (1852), 4 (6): 290-300 (1852), 4 (7): 324-343 (1852), 4 (9): 422-432, pl. 10 (1852), 5 (7): 310-322 (1853)
MacLeay, [1826]
in King, Narrative of a Survey of the Intertropical and Western Coasts of Australia. Catalogue of Insects, collected by Captain King, R.N. in King, Narr. Surv. Austr. 2 : 438-469
Moore, 1872
Descriptions of new Indian Lepidoptera Proc. zool. Soc. Lond. 1872 (2) : 555-583, pl. 32-34
Moore, [1881]
The Lepidoptera of Ceylon Lepid. Ceylon 1 (1): 1-40, pl. 1-18 (Dec 1880), (2): 41-80, pl. 19-36 (Jan 1881), (3): 81-136, pl. 37-54 (Jun 1881), (4): 137-190, pl. 55-71 (Dec 1881)
Oberthür, 1878
Faune des Lépidoptères de la côte orientale d'Afrique Étud. d'Ent. 3 : 11-36, pl. 1-3
Piepers & Snellen, 1909
The Rhopalocera of Java. Pieridae Rhop. Java [1] : 1-65, pl. 1-4
Rebel, 1914
Wissenschaftliche Ergebnisse der Expedition R. Grauernach. Zentralafrika. 1909-1911. Lepidoptera Ann. Mus. Wien. 28 : 219-294, pl. 17-24
Rogenhofer, 1889
Afrikanische Schmetterling des k. k. naturhist. Hofmuseums Ann. Mus. Wien. 4 : 547-554, pl. 23
Sharpe, 1891
Descriptions of New Butterflies collected by Mr. F. J. Jackson, F.Z.S:, in British East Africa, during his recent Expedition. - Part I & II Proc. Zool. Soc. Lond. 1891 : 187-194, pl. 16-17, : 633-638, pl. 48
Sheljuzhko, 1908
Quelques formes nouvelles des Lépidopteres Rev. russ. ent. 7 (4) : 232-234
[EBW]; Smart, 1976
The Illustrated Encyclopedia of the Butterfly World Illust. Encyp. Butt. World: -
Linnaeus in Sparrman, 1768
Iter in Chinam quod praeside D. D. Car. v. Linné Amoenitates Acad. 7 : 497-506
Stoll, [1781]
Uitlandsche Kapellen (Papillons exotiques) in Cramer, Uitl. Kapellen 4 (26b-28): 29-90, pl. 305-336 (1780), (29-31): 91-164, pl. 337-372 (1781), (32-32): 165-252, 1-29, pl. 373-400 (1782)
Stoneham, 1940
Notes on the African genus Colotis, Hubnr. (Teracolus Swin.) and some new forms Bull. Stoneham Mus. (40): 1-4
Suffert, 1904
Neue afrikansche Tagfalter aus dem kön. zool. Museum, Berlin, und meiner Sammlung Dt. Ent. Z. Iris 17 (1) : 12-107, pl. 1-3
Suffert, 1904
Neue Tagfalter aus Deutsch-Ost-Afrika Dt. Ent. Z. Iris 17 (1) : 124-132
Talbot, 1928
A new Pierine from Kenya Colony Bull. Hill. Mus. 2 : 29
Talbot, 1943
Revisional notes on Anapheis Hubn. and Belenois Hubn., with descriptions of some new forms (Lep. Rhop. Pieridae) Ann. Mag. nat. Hist. (11) 10 (65) : 305-336
[BMAT]; Tennent, 1996
The Butterflies of Morocco, Algeria and Tunisia : 1-217
Trimen, 1883
Descriptions of twelve new species of South African Lepidoptera Rhopalocera Trans. ent. Soc. Lond. 1883 (4) : 347-363
[BRU]; Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997
Guide to the Butterflies of Russia and adjacent territories: Hesperiidae, Papilionidae, Pieridae, Satyridae Butts. Russia adj. terr. 1
Vane-Wright & de Jong, 2003
The butterflies of Sulawesi: annotated checklist for a critical island faunda Zool. Verh. Leiden 343: 3-268, pl. 1-16
Vollenhoven, 1869
Description des espèces nouvelles de coléoptères, orthoptères, lépidoptères et hémiptères, mentionees dans da liste précédente Faune Madag. 5 (Ins.) : 7-24, pl. 1-2
Wallengren, 1863
Lepidopterologische Mittheilungen (1-3) Wien ent. Monats. 4 (2): 33-46, 4 (6): 161-176 (1860), 7 (5): 137-151 (1863)
Ward, 1870
Descriptions of new species of diurnal Lepidoptera from Madagascar Ent. mon. Mag. 7 : 30-32
Winhard, 2000
Pieridae I Butterflies of the world 10 : 1-40, pl. title,1-48,back
[BIR]; Wynter-Blyth, 1957
Butterflies of the Indian Region (1982 Reprint)

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