500x358 (~45Kb) upperside Plymbridge woods, Devon, UK 15/08/96, Photo © Simon Coombes | 450x332 (~41Kb) underside Plymbridge woods, Devon, UK 19/07/93, Photo © Simon Coombes | 600x410 (~45Kb) upperside Finland, Joutseno, 7/1995, Photo © Tero Piirainen |
500x427 (~37Kb) underside Finland, Joutseno, 7/1995, Photo © Tero Piirainen | 600x606 (~45Kb) ♂ upperside FINLAND: Ka: Miehikkälä, 673:53-54, 18.7.1997, Photo © Markku Savela | 533x400 (~48Kb) ♂ ♀ underside A pine forest edge, Akademgorodok [Academy Town], Novosibirsk, West Siberia, Russia. 11th July 1995, Photo © Oleg Kosterin |
512x768 (~80Kb) ♂ upperside Germany, Baden-Württemberg, Horrheim, 8° 59' E, 48° 59' N, altitude 350m, 12.07.1997 (sucking on Cirsium arvense), Photo © Karl Hofsäß | 621x414 (~48Kb) A long forb meadow at the Adagum River right bank, the town Krymsk, Krymsk District, Krasnodarskii Krai Province, S Russia. 27th June 1991 (A small congregation on a large inflorescence of Heracleum dissectum), Photo © Oleg Kosterin | 595x400 (~75Kb) larva Central Europe 1990-2000, Photo © Michael Borsch |
594x400 (~58Kb) ♂ upperside Central Europe 1990-2000, Photo © Michael Borsch | 400x598 (~40Kb) pupa Central Europe 1990-2000, Photo © Michael Borsch | 600x800 (~64Kb) underside Sweden, Ekebo, Julita, 26.7.2002, Photo © Leif Wahlberg |
1196x897 (~133Kb) ♀ upperside Sweden, Ekebo, Julita, 12.8.2004, Photo © Leif Wahlberg | 806x1075 (~158Kb) Sweden, Ekebo, Julita, 5.8.2004, Photo © Leif Wahlberg | 470x435 (~96Kb) ♀ upperside Czech Republic, Blansko, 6.8.2004, Photo © Michal Koupý |
470x455 (~93Kb) underside Czech Republic, Blansko, 31.7.2004, Photo © Michal Koupý | 850x645 (~101Kb) ♀ upperside Russia, Moscow area, 22.7.2008, Photo © D. Smirnov | 548x633 (~111Kb) ♀ underside Russia, Moscow area, 29.7.2008, Photo © D. Smirnov |
1088x341 (~103Kb) larva Russia, Moscow area, 5.6.2011 (36°25'E, 56°00'N), Photo © D. Smirnov | 575x677 (~111Kb) ♂ upperside Russia, Moscow area, 29.7.2008, Photo © D. Smirnov | 1066x788 (~238Kb) ♂ Lithuania: Dzukija, Trakiskiaei, 15.7.2008, Photo © Harri Arkkio |
930x729 (~97Kb) underside Lithuania: Dzukija, Marcinkonys-Zervynos, 15.7.2008, Photo © Harri Arkkio |
Larva on Viola palustris?, V. canina? [SPRK], Violaceae , Rubus spp. [MRS]
Larva on Viola mumbyana?, (in Spain) V. odorata, V. canina [BMAT]
Larva on Viola variegata [BRU2]: 34
951x624 (~42Kb) ♀ upperside Oppadake, Okinawa, Ryukyus, Japan, 9-93, Photo © S. Shuichi Haupt | 951x624 (~46Kb) ♂ underside Funaura, Iriomote, Ryukyus, Japan, 5-95, Photo © S. Shuichi Haupt | 658x514 (~114Kb) upperside An opening in a Cryptomeria forest at the top of Kongo Mt., Temporin temple complex, the border of Osaka and Nara Prefectures, Honshu, Japan, 3rd August 2002, Photo © Oleg Kosterin |
765x634 (~117Kb) underside An opening in a Cryptomeria forest at the top of Kongo Mt., Temporin temple complex, the border of Osaka and Nara Prefectures, Honshu, Japan, 3rd August 2002, Photo © Oleg Kosterin | 1600x1200 (~188Kb) ♀ upperside Japan: Tokyo, 23.8.2007, Photo © Peter Payzant | 1600x1200 (~181Kb) ♂ upperside Japan: Tokyo, 24.8.2007, Photo © Peter Payzant |
512x341 (~52Kb) India: Ziro, 9.11.2005, Photo © Kedar Tokekar | 512x341 (~48Kb) upperside India: Ziro, 9.11.2005, Photo © Kedar Tokekar |
Larva on Viola caespitosa, V. yedoensis, V. betonicifolia, V. formosana, V. nagasawai, V. philippica, V. tricolor, V. verecunda [MRS], Viola betonicifolia, V. hederacea, V. odorata Dunn & Dunn, 1991, Review Austr. Butts.
748x560 (~72Kb) underside Montejo de la Sierra, prov. Madrid, Spain, July-1988, Photo © Enrique Garcia-Barros | 554x596 (~96Kb) upperside Port-Krym, Crimea, Ukraine. 10th June 1990, Photo © Oleg Kosterin | 1076x956 (~337Kb) underside Iran, Ostan-e Lorestan, Shakhrestan-e Borujerd , the Hayan village environs, Gavimir Chashmah. 25th May 2017, Photo © Oleg Kosterin |
1198x1021 (~516Kb) underside Iran, Ostan-e Lorestan, Khorramabad-e Shakhrestan, the Sezar River right bank 2 km downstream Bisheh Waterfall. 26th May 2017, Photo © Oleg Kosterin | 1442x1330 (~720Kb) underside Iran, Ostan-e Lorestan, Khorramabad-e Shakhrestan, the Sezar River right bank 2 km downstream Bisheh Waterfall. 26th May 2017, Photo © Oleg Kosterin | 1003x977 (~555Kb) underside Iran, Ostan-e Lorestan, Khorramabad-e Shakhrestan, the Sezar River right bank 2 km downstream Bisheh Waterfall. 26th May 2017, Photo © Oleg Kosterin |
Larva on Viola , V. tricolor [H&R], (in Israel) Viola modesta [BMAT]
600x435 (~32Kb) upperside Morocco, Azrou, Moyen Atlas central, Photo © Michel Tarrier | 600x435 (~31Kb) underside Morocco, Forêt de Jaâba, Moyen Atlas central, Photo © Michel Tarrier |
Larva on Viola grypoceras, V. verecunda, V. eizanensis [MRS], Viola uniflora [BRU2]: 35
533x400 (~47Kb) ♀ upperside An open larch (Larix gmelinii (Rupr.) Rupr.)/birch (Betula pendula Roth. and B. davurica Pall.) forest above the left bluff of the Budyumkan River valley 5 km of its confluence with the Argun' River 6 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 2nd August 1997, Photo © Oleg Kosterin | 800x600 (~97Kb) ♀ underside An open larch/birch stand on a southern slope of the Polovinnaya Pad' valley few kilometres of the Argun' River left bank 13 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 28th July 1997, Photo © Oleg Kosterin |
800x800 (~146Kb) ♀ upperside A woody Bol'shoi Tesh River right bank 7 km NEE of the village Kuzedeevo, the piedmonts of Gornaya Shoria Mts., Novokuznetsk District., Kemerovo Province, West Siberia, Russia. 27th July 1996, Photo © Oleg Kosterin | 514x403 (~26Kb) ♂ upperside Tall herbage on a woody Bol'shoi Tesh River right bank 7 km NEE of the village Kuzedeevo, the piedmonts of Gornaya Shoria Mts., Novokuznetsk District., Kemerovo Province, West Siberia, Russia. 27th July 1996, Photo © Oleg Kosterin | 533x400 (~40Kb) ♂ underside Tall herbage on a woody Bol'shoi Tesh River right bank 7 km NEE of the village Kuzedeevo, the piedmonts of Gornaya Shoria Mts., Novokuznetsk District., Kemerovo Province, West Siberia, Russia. 27th July 1996, Photo © Oleg Kosterin |
533x400 (~57Kb) A southern slope, overgrown with bushes of Spiraea and Caragana arborescens, of the valley of a brook being a left tributary of the Bol'shoi Tesh River 2 km upstream of its junction with the Malyi Tesh, within the famous "lime-tree island" of the forests of a relic species Tilia sibirica Fisch., 10 km east of the village Kuzedeevo, the elevation of Gornaya Shoriya, Novokuznetsk District, Kemerovo Province, West Siberia, Russia. 12th July 1995, Photo © Oleg Kosterin | 300x400 (~19Kb) 3th June 1995. The caterpillar was collected on a southern slope, overgrown with bushes of Spiraea and Caragana arborescens, of the valley of a brook being a left tributary of the Bol'shoi Tesh River 2 km upstream of its junction with the Malyi Tesh, within the famous "lime-tree island" of the forests of a relic species Tilia sibirica Fisch., 10 km east of the village Kuzedeevo, the elevation of Gornaya Shoriya, Novokuznetsk District, Kemerovo Province, West Siberia, Russia. 12th July 1995, Photo © Oleg Kosterin | 800x961 (~139Kb) 6th June 1995. The caterpillar was collected on a southern slope, overgrown with bushes of Spiraea and Caragana arborescens, of the valley of a brook being a left tributary of the Bol'shoi Tesh River 2 km upstream of its junction with the Malyi Tesh, within the famous "lime-tree island" of the forests of a relic species Tilia sibirica Fisch., 10 km east of the village Kuzedeevo, the elevation of Gornaya Shoriya, Novokuznetsk District, Kemerovo Province, West Siberia, Russia. 21st May 1995, Photo © Oleg Kosterin |
400x533 (~38Kb) 6th June 1995. The caterpillar was collected on a southern slope, overgrown with bushes of Spiraea and Caragana arborescens, of the valley of a brook being a left tributary of the Bol'shoi Tesh River 2 km upstream of its junction with the Malyi Tesh, within the famous "lime-tree island" of the forests of a relic species Tilia sibirica Fisch., 10 km east of the village Kuzedeevo, the elevation of Gornaya Shoriya, Novokuznetsk District, Kemerovo Province, West Siberia, Russia. 21st May 1995, Photo © Oleg Kosterin | 400x533 (~34Kb) 6th June 1995. The caterpillar was collected on a southern slope, overgrown with bushes of Spiraea and Caragana arborescens, of the valley of a brook being a left tributary of the Bol'shoi Tesh River 2 km upstream of its junction with the Malyi Tesh, within the famous "lime-tree island" of the forests of a relic species Tilia sibirica Fisch., 10 km east of the village Kuzedeevo, the elevation of Gornaya Shoriya, Novokuznetsk District, Kemerovo Province, West Siberia, Russia. 21st May 1995, Photo © Oleg Kosterin | 400x533 (~38Kb) 6th June 1995. The caterpillar was collected on a southern slope, overgrown with bushes of Spiraea and Caragana arborescens, of the valley of a brook being a left tributary of the Bol'shoi Tesh River 2 km upstream of its junction with the Malyi Tesh, within the famous "lime-tree island" of the forests of a relic species Tilia sibirica Fisch., 10 km east of the village Kuzedeevo, the elevation of Gornaya Shoriya, Novokuznetsk District, Kemerovo Province, West Siberia, Russia. 21st May 1995, Photo © Oleg Kosterin |
400x533 (~36Kb) 6th June 1995. The caterpillar was collected on a southern slope, overgrown with bushes of Spiraea and Caragana arborescens, of the valley of a brook being a left tributary of the Bol'shoi Tesh River 2 km upstream of its junction with the Malyi Tesh, within the famous "lime-tree island" of the forests of a relic species Tilia sibirica Fisch., 10 km east of the village Kuzedeevo, the elevation of Gornaya Shoriya, Novokuznetsk District, Kemerovo Province, West Siberia, Russia. 21st May 1995, Photo © Oleg Kosterin | 800x600 (~108Kb) 6th June 1995. The caterpillar was collected on a southern slope, overgrown with bushes of Spiraea and Caragana arborescens, of the valley of a brook being a left tributary of the Bol'shoi Tesh River 2 km upstream of its junction with the Malyi Tesh, within the famous "lime-tree island" of the forests of a relic species Tilia sibirica Fisch., 10 km east of the village Kuzedeevo, the elevation of Gornaya Shoriya, Novokuznetsk District, Kemerovo Province, West Siberia, Russia. 12th July 1995, Photo © Oleg Kosterin | 900x600 (~136Kb) ♂ upperside A road in a pine forest, the Stolby Nature Reserve, Krasnoyarsk Province, 11th July 1995 (on Rubus idaeus), Photo © Oleg Kosterin |
Larva on Viola , V. palustris [H&R], Viola grypoceras, Filipendula camtschatica [MRS]
24.4.2023 (10)
If you have corrections, comments or
information to add into these pages, just send mail to
Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.