Life Insecta Lepidoptera Ditrysia Hesperioidea Hesperiidae Heteropterinae

• Carterocephalus palaemon (Pallas, 1771) EU, AN, AC, Japan. See [maps] Arctic Skipperling Schacksmygare Gulfläckig glanssmygare Chequered Skipper Kosternik palemon Keltatäplähiipijä L'Echiquier Arctic Skipper Gelbwürfeliger Dickkopffalter

  • Papilio palaemon Pallas, 1771; Reise Russ. Reich. 1: 471
  • Papilio paniscus Fabricius, 1775; Syst. Ent.: 531
  • brontes Denis & Schiffermüller, 1775
  • Pamphila palaemon borealis Lingonblad, 1945; Notulae Ent. 24 (3-4): 74; TL: N.Finland

  • Carterocephalus palaemon mackenziei; [LepIndex]; [CoL]
  • Carterocephalus palaemon; [SPRK]; [H&R], 332; [BOW]: pl. 11, f. 12; [BRU], 124; [NACL], #3982; [NAB], 771; [MRS], 724; [BOC], 66; [Opler]; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 12; [RFEL]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [BAMONA] ; [SVF]
  • Pamphila palaemon; Dyar, 1903, Bull. U.S. nat. Mus. 52: 48

  700x584 (~61Kb) upperside Finland, Sotkamo, Lontanniemi 712:54, 21.6.1997, Photo © Tero Piirainen	600x580 (~39Kb) underside Finland, Sotkamo, Kontinjoki 711:54, 21.6.1997, Photo © Tero Piirainen	700x542 (~49Kb) upperside USA: Near Crystal Mtn, (46°57'51N 121°28'55W), Pierce Co., WA, 27.7.1999, Photo © Markku Savela
  478x604 (~52Kb) underside Germany, Baden-Württemberg, Maulbronn, Waldgebiet S Köpfle (8° 50' E, 49° 01' N, 300m), 13.05.2000 (on Lychnis flos-cuculi), Photo © Karl Hofsäß	1292x1280 (~179Kb) underside Canada, British Columbia, Brilliant, 1.5.2005, Photo © Norbert Kondla	646x616 (~87Kb) Russia, Moscow area (36°25'E, 56°00'N), 15.6.2009, Photo © D. Smirnov
  786x647 (~110Kb) ♀ Russia, Moscow area (36°25'E, 56°00'N), 15.6.2009, Photo © D. Smirnov	744x601 (~101Kb) ♂ Russia, Moscow area (36°25'E, 56°00'N), 14.6.2009, Photo © D. Smirnov	810x640 (~132Kb) underside Russia, Khabarovskiy Kray, Khabarovsk southern env., Bychikha village. 7th July 2014, Photo © Oleg Kosterin
  1072x810 (~192Kb) Russia, Khabarovskiy Kray, Khabarovsk southern env., Bychikha village. 7th July 2014, Photo © Oleg Kosterin	879x871 (~159Kb) Russia, Khabarovskiy Kray, Khabarovsk southern env., Bychikha village. 7th July 2014, Photo © Oleg Kosterin

  Larva on Calamagrostis purpurea? [SPRK], Bromus , Cynosurus cristatus, Brachypodium , Calamagrostis [BRU], Calamagrostis purpurascens [NAB]

  • C. p. palaemon

    • Carterocephalus palaemon palaemon; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 12
  • C. p. albiguttata Christoph, 1893 Ural - Siberia - Far East

    • Carterocephalus palaemon var. albiguttata Christoph, 1893; Dt. Ent. Z. Iris 6 (1): 87
    • australis Kurenzov, 1941

    • Carterocephalus palaemon albiguttata; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 12; Yakovlev, 2012, Nota lepid. 35 (1): 61; [LepIndex]; [CoL]
    • Carterocephalus palaemon albiguttatus; [MRS], 724

    502x365 (~18Kb) A bushy meadow on the second right bank terrace of the Koen River at the mouth of the Volchikha Rivulet, just downstream of the village Nizhnii Koen, Iskitim District, Novosibirsk Province, West Siberia, Russia. 6th June 1993 (C. silvicolus wings closed at left), Photo © Oleg Kosterin

    472x720 (~80Kb) upperside A valley meadow in a birch forest in the Shadrikha Rivulet valley 2 km upstream of the village Melnichikha, Novosibirsk District and Province, West Siberia, Russia. 27th May 1995 (with C. silvicola below), Photo © Oleg Kosterin

    533x400 (~39Kb) ♂ underside An open birch forest in the Zyryanka Rivulet valley near its fall into the Novosibirsk Water Reserve. 29th May 1997, on Geranium sylvaticum L. Photo © Oleg Kosterin

  • C. p. satakei (Matsumura, 1919) Japan

    • palaemon satakei (Matsumura, 1919) (Pamphila); Thous. Ins. Japan. Addit. 3: 669

    • Carterocephalus palaemon satakei; Kudrna, 1974, Atalanta 5: 117; [LepIndex]; [CoL]
  • C. p. murasei (Matsumura, 1925) Sakhalin

    • Pamphila palaemon murasei Matsumura, 1925; J. Coll. Agric. Hokkaido imp. Univ. 15: 107, pl. 11, f. 14 ♂; TL: Saghalien
  • C. p. tolli Krzywicki, 1967

    • Carterocephalus palaemon tolli Krzywicki, 1967; Ann. zool. Warsz. 25 (1): (1-213)

    • Carterocephalus palaemon tolli; [LepIndex]; [CoL]
  • C. p. akaishianus Fujioka, 1970 Japan

    • Carterocephalus palaemon akaishianus; [LepIndex]; [CoL]
  • C. p. taigana (Kurenzov, 1970)

    • palaemon taigana (Kurenzov, 1970) (Pamphila); 364

    • Carterocephalus palaemon taigana; Korb & Bolshakov, 2011, Eversmannia Suppl. 2:
  • C. p. magnus Tilden & Emmel, 1998 ?, British Columbia - California

    • Carterocephalus palaemon magnus Tilden & Emmel, 1998; ; TL: Plantation, Sonoma Co., California

    • Carterocephalus palaemon magnus; [BOC], 66
    • Carterocephalus skada magnus; [P&N23]; [iNaturalist]
• Carterocephalus mandan (Edwards, 1863) Canada. See [maps]

  • Hesperia mandan Edwards, 1863; Proc. ent. Soc. Philad. 2 (1): 20; TL: Pine Ridge, Manitoba

  • Carterocephalus mandan mandan; [P&N23]; [iNaturalist]
  • Carterocephalus mandan; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 30; [MPG] ; [P&N23]; [iNaturalist] ; [LEPIFORUM] ; [BOA]
  • Carterocephalus palaemon mandan; [NACL], #3982a; [BOC], 66; [LepIndex]; [CoL]
  • C. m. mesapano (Scudder, 1868) New England

    • Hesperia mesapano Scudder, 1868; Proc. Boston Soc. Nat. Hist. 11: 383; TL: Norway, New England

    • Carterocephalus mandan mesapano; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 30; [P&N23]; [iNaturalist] ; [BOA]
    • Carterocephalus palaemon mesapano;
• Carterocephalus skada (Edwards, 1870) Cascades? - Alaska. See [maps]

  • skada Edwards, 1870; ; TL: Kodiak, Alaska
  • skada Edwards, 1871; (preocc. Edwards, 1870)

  • Carterocephalus palaemon skada; [BOC], 66
  • Carterocephalus skada skada; [P&N23]; [iNaturalist]
  • Carterocephalus skada; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 30; [MPG] ; [P&N23]; [iNaturalist] ; [LEPIFORUM] ; [BOA] ; [BOA]

  600x520 (~53Kb) upperside USA: Bridge Creek Trailhead, North Cascades Highway, (48°30'30N 120°43'00W), Chelan Co., WA, 28.7.1999, Photo © Markku Savela

  668x581 (~50Kb) USA: Washington, Snohomish Co., Kelcema Lake, 3.8.2011, Photo © Markku Savela

• Carterocephalus silvicola (Meigen, 1829) Fennoscandia, Siberia, Amurland, Kamschatka, Japan. See [maps] Svartfläckig glanssmygare Northern Checquered Skipper Fläcksmygare Kosternik leśniak Mustatäplähiipijä

  • Hesperia silvicola Meigen, 1829; Syst. Beschr. eur. Schmett. 2: 65, pl. 55, f. 7A-C
  • silvius (Knoch, 1781) (Papilio); Beitr. Insektengesch. 1: pl. 5
  • Pamphila silvius isshikii Matsumura, 1925; J. Coll. Agric. Hokkaido imp. Univ. 15: 107, pl. 8, f. 17 ♂; TL: Saghalin
  • Pamphila silvius isshikii ab. doii Matsumura, 1928; Insecta Matsumurana 2 (4): 201; TL: Kuriles
  • Pamphila silvius shikotanus Nakahara, 1926; Ins. Inscit. Mens. 14: 51; TL: Kuriles

  • Carterocephalus silvicola malaisei; [iNaturalist]
  • Carterocephalus silvicola trybomi; [iNaturalist]
  • Carterocephalus silvicola virilis; [iNaturalist]
  • Carterocephalus silvicola; [SPRK]; [BRU], 124; [MRS], 724; Yakovlev, 2012, Nota lepid. 35 (1): 62; [RFEL]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]
  • Carterocephalus silvicolus; [H&R], 333; [BOW]: pl. 11, f. 13; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 12; [MBE] ; [SVF]
  • Carterocephalus sylvicola isshikii; Kudrna, 1974, Atalanta 5: 117
  • Pamphila silvius isshikii; Matsumura, 1926, Insecta Matsumurana 1 (2): 106
  • sylvius[sic];
  • ?sylvicola[sic];

  600x374 (~29Kb) FINLAND: Ab: Suomusjärvi, 669:31, 21.6.1996, Photo © Markku Savela	900x700 (~129Kb) ♂ underside FINLAND, Ab: Suomusjärvi, 669:31, 28.5.2000, Photo © Markku Savela	600x533 (~39Kb) FINLAND: Ab: Suomusjärvi, 669:31, 21.6.1996, Photo © Markku Savela
  500x537 (~40Kb) ♀ upperside Finland, Lempäälä, Koipijärvi, 6816:331, 24.6.1996, Photo © Tero Piirainen	520x600 (~36Kb) ♂ upperside Finland, Kajaani, Kiviranta 711:54, 21.6.1997, Photo © Tero Piirainen	497x750 (~39Kb) ♂ upperside Sipoo 670:39, 07.06.1981, Photo © Pertti Pakkanen
  464x500 (~32Kb) ♀ upperside Alpinotype meadow patches among sparce dwarf alder growth 3 km SE of the top or the mountain Yagodnaya, the Khalzan mountain range, 600 m above sea level, 55 km SW of Petropavlovsk- Kamchatskii, S Kamchatka, Russia. 16th August 1992 (on Saussuraea), Photo © Oleg Kosterin	500x304 (~15Kb) ♂ upperside Open birch/pine forest in the Zyryanka Rivulet valley, Akademgorodok [Academy Town], Novosibirsk, West Siberia, Russia. 8th June 1998, Photo © Oleg Kosterin	795x596 (~98Kb) ♀ Russia, Moscow area (36°25'E, 56°00'N), 14.6.2009, Photo © D. Smirnov
  728x602 (~101Kb) ♂ Russia, Moscow area (36°25'E, 56°00'N), 1.6.2009, Photo © D. Smirnov	1135x932 (~282Kb) ♂ upperside Russia, West Siberia, the Altai Mts., Altai Republic, Ust'-Koksa District, the Katunskii mountain range northern principal slope, the eastern slope of the Kuzuyak Pass through a ridge dividing the Kucherla and Akkem River valleys (the Kucherla side), a muddy road through birch/larch forest, about 1200 m above sea level. 26th June 2007, Photo © Oleg Kosterin

  Larva on Gramineae , Calamagrostis purpurea, Elytrigia repens [SPRK], Bromus , Cynosurus , Milium effusum [BRU]

• Carterocephalus argyrostigma (Eversmann, 1851) S.Siberia, Mongolia, NE.China. See [maps]

  • Hesperia argyrostigma Eversmann, 1851; Bull. Soc. imp. Nat. Moscou 24 (2): 624; TL: Irkutsk, Kyakhta, Transbaikalia
  • Cyclopides argenteogutta Butler, 1870; Trans. ent. Soc. Lond. 1870 (4): 512; TL: Nubia

  • Carterocephalus argyrostigma; [BOW]: pl. 207, f. 23; [BRU], 124; Yakovlev, 2012, Nota lepid. 35 (1): 61; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 12; [RFEL]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]
  • Hesperia argyrostigma; Nordmann, 1851, Bull. Soc. imp. Nat. Moscou 24: 442, pl. 12, f. 1-2

  800x600 (~76Kb) upperside A meadow steppe, at granite rocks, on a moderate elevation Adon-Chelon, 10 km NNW of the village Tasyrkhoi, SE Chita Province (Dahuria), Transbaikalia, Siberia, Russia. 21st June 1995, Photo © Oleg Kosterin

  533x400 (~34Kb) underside A meadow steppe, at granite rocks, on a moderate elevation Adon-Chelon, 10 km NNW of the village Tasyrkhoi, SE Chita Province (Dahuria), Transbaikalia, Siberia, Russia. 21st June 1995, Photo © Oleg Kosterin

  1000x1000 (~152Kb) upperside A meadow steppe, at granite rocks, on a moderate elevation Adon-Chelon, 10 km NNW of the village Tasyrkhoi, SE Chita Province (Dahuria), Transbaikalia, Siberia, Russia. 21st June 1995, Photo © Oleg Kosterin

• Carterocephalus dieckmanni Graeser, 1888 NE.China - S.China, Amur, Ussuri - N.Burma. See [maps]

  • Carterocephalus dieckmanni Graeser, 1888; Berl. Ent. Zs. 32 (1): 102; TL: Vladivostok, Ussuri region

  • Carterocephalus dieckmanni watsoni; [LepIndex]; [CoL]
  • Carterocephalus dieckmanni; [BOW]: pl. 207, f. 28 (text); [BRU], 125; [MRS], 724; [RFEL]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [SZMN]
  • Carterocephalus diekmanni[sic, dieckmanni]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 11
  • C. d. gemmatus Leech, 1891

    • Carterocephalus gemmatus Leech, 1891; Entomologist 24 (Suppl.): 59; TL: Ta-chein-lu, Put-tsu-fong; Wa-ssu-kou

    • Carterocephalus dieckmanni gemmatus; [MRS], 724; [LepIndex]; [CoL]
    • Carterocephalus gemmatus; [iNaturalist]
  • C. d. minor (Evans)

    • Carterocephalus dieckmanni minor; [MRS], 724; [LepIndex]; [CoL]
  • C. d. soyosanus Murayama, 1983

    • Carterocephalus dieckmanni soyosanus Murayama, 1983; Entomotaxonomia 5: (281-288)
• Carterocephalus christophi Grum-Grshimailo, 1891 Qinghai, W.Sichuan, Yunnan. See [maps]

  • Carterocephalus christophi Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4): 460; TL: montibus Sinin-Schan
  • Aubertia dulcis Oberthür, 1896; Étud. d'Ent. 20: 40, pl. 9, f. 16♂

  • Carterocephalus christophi christophi; [iNaturalist]
  • Carterocephalus christophi; [BOW]: pl. 207, 28 (text); [MRS], 724; Huang, 2019, Neue Ent. Nachr. 78: 215; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]
• Carterocephalus tibetanus (South, 1913) SE.Tibet. See [maps]

  • Aubertia niveomaculatus tibetanus South, 1913; J. Bomb. nat. Hist. Soc. 22 (3): 612
  • Carterocephalus chayuensis Lee, 1979; 36, pl. 1, f. 5-6; TL: Nanxue, Chayu
    • = ; Huang, 2019, Neue Ent. Nachr. 78: 216

  • Carterocephalus christophi tibetanus; [MRS], 724; [LepIndex]; [CoL] ; [iNaturalist]
  • Carterocephalus tibetanus; Huang, 2019, Neue Ent. Nachr. 78: 216
• Carterocephalus alcina Evans, 1939 Yunnan. See [maps]

  • Carterocephalus alcina Evans, 1939; Proc. R. ent. Soc. Lond. (B) 8 (8): 164 [?]

  • Carterocephalus alcina; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Yutaka Inayoshi]
  • Carterocephalus alcinus; ; [SZMN]
• Carterocephalus alcinoides Lee, 1962 Yunnan. See [maps]

  • Carterocephalus alcinoides Lee, 1962; Acta ent. sin. 11 (2): 141, 146

  • Carterocephalus alcinoides; [MRS], 724; [LepIndex] ; [CoL] ; [iNaturalist]

• Carterocephalus avanti (de Nicéville, 1886) Tibet, W.China. See [maps]

  • Pamphila avanti de Nicéville, 1886; J. Asiat. Soc. Bengal 55 Pt.II (3): 255, pl. 11, f. 10 ♂; TL: Sikkim

  • Carterocephalus avanti avanti; [iNaturalist]
  • Carterocephalus avanti ensis; [LepIndex]; [CoL] ; [iNaturalist]
  • Carterocephalus avanti flavostigma; [CoL]
  • Carterocephalus avanti; [BOW]: pl. 207, f. 24; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]
• Carterocephalus canopunctatus Nabokov, 1941 Sichuan. See [maps]

  • Carterocephalus canopunctatus Nabokov, 1941; 222; TL: Ta-Tsien-Lu

  • Carterocephalus canopunctatus; Huang, 2019, Neue Ent. Nachr. 78: 216
• Carterocephalus flavomaculatus Oberthür, 1886 W.China, Tibet. See [maps]

  • Carterocephalus flavomaculatus Oberthür, 1886; Étud. d'Ent. 11: 27

  • Carterocephalus flavomaculatus; [BOW]: pl. 207, f. 28 (text); [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]
• Carterocephalus houangty Oberthür, 1886 W.China. See [maps]

  • Carterocephalus houangty Oberthür, 1886; Étud. d'Ent. 11: 27

  • Carterocephalus houangty bootia; [LepIndex]; [CoL] ; [iNaturalist]
  • Carterocephalus houangty; [BOW]: pl. 207, f. 25; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]
  • C. h. shoka Evans, 1915 SE.Tibet

    • Carterocephalus houangty shoka Evans, 1915; J. Bombay nat. Hist. Soc. 23 (3): 546

    • Carterocephalus houangty shoka; Huang, 2001; [LepIndex]; [CoL] ; [iNaturalist]
  • C. h. jiuzaikouensis Yoshino, 2001 N.Sichuan

    • Carterocephalus houangty jiuzaikouensis Yoshino, 2001; Futao (38): 12, pl. 3, f. 5-6, 9-10; TL: Jiuzaikou Valley, Nanping, SE. Minshan, N.Sichuan

    • Carterocephalus houangty jiuzaikouensis; Yoshino, 2003, Futao (43): 10 (note)
  • C. h. zorgensis Yoshino, 2003 N.Sichuan

    • Carterocephalus houangty zorgensis Yoshino, 2003; Futao (43): 10, f. 33-34; TL: 70km NW of Songpan town, 3500m, SE.Minshan, N. Sichuan
• Carterocephalus micio Oberthür, 1891 China. See [maps]

  • Carterocephalus micio Oberthür, 1891; Étud. d'Ent. 15: 19

  • Carterocephalus micio; [BOW]: pl. 207, f. 26; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]
• Carterocephalus niveomaculatus Oberthür, 1886 W.Sichuan, N.Yunnan. See [maps]

  • Carterocephalus niveomaculatus Oberthür, 1886; Étud. d'Ent. 11: 27; TL: Ta-Tsien-Lou

  • Carterocephalus niveomaculatus; [BOW]: pl. 207, f. 28; Huang, 2019, Neue Ent. Nachr. 78: 215; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]
• Carterocephalus pulchra (Leech, 1891) Tibet, W.China. See [maps]

  • Pamphila pulchra Leech, 1891; Entomologist 24 (Suppl.): 59; TL: Ta-chein-lu; Wa-ssu-kou
  • Carterocephalus ops Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4): 460

  • Carterocephalus pulchra ops; [LepIndex]; [CoL] ; [iNaturalist]
  • Carterocephalus pulchra pulchra; [iNaturalist]
  • Carterocephalus pulchra; [BOW]: pl. 207, f. 27; [LepIndex] ; [CoL] ; [iNaturalist]
• Carterocephalus abax Oberthür, 1886 E.Tibet. See [maps]

  • Carterocephalus abax Oberthür, 1886; Étud. d'Ent. 11: 27, pl. 5, f. 27

  • Carterocephalus abax abax; [iNaturalist]
  • Carterocephalus abax patra; [LepIndex]; [CoL] ; [iNaturalist]
  • Carterocephalus abax; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]
• Carterocephalus habaensis Yoshino, 1997 Yunnan. See [maps]

  • Carteocepharus[sic] habaensis Yoshino, 1997; Neo Lepidoptera 2: 8, f. 54-55, 64; TL: Zhondian, 3000m, N. Yunnan
• Carterocephalus zhongkui Sakai, 2016 Gansu. See [maps]

  • Carterocephalus niveomaculatus zhongkui Sakai, 2016; 1; TL: Xiahe, Gansu

  • Carterocephalus zhongkui; Huang, 2019, Neue Ent. Nachr. 78: 216
• Carterocephalus bozanoi Huang, 2019 Xizang. See [maps]

  • Carterocephalus bozanoi Huang, 2019; Neue Ent. Nachr. 78: 216; TL: China, Xizang, Gongbujiangda Co., 30km W of Gongbujiangda Town, 3650m

26.3.2023 (21)

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.