Hesperiinae

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All	Heteropterinae	Trapezitinae	Hesperiinae	(hesperiinae)	Grass Skippers
			Hesperiinae	(hesperiinae)
Photo	Zariaspes	Hesperiidae			Grass Skippers
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Hesperiinae Latreille, 1809 Monocot Skippers Folded-winged Skippers Grass Skippers

Hesperiinae; [CoL] ; [iNaturalist]
Hesperiinae (Hesperiidae); [BK], 408; [NL4A], 61; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 11; [RFEL]

Synapte Mabille, 1904 — 7 spp.
Corticea Evans, 1955 — 12 spp.
Callimormus Scudder, 1872 — 9 spp.
Radiatus Mielke, 1968 — 1 sp.
Peba Mielke, 1968 — 1 sp.
Vidius Evans, 1955 — 15 spp.
Monca Evans, 1955 — 4 spp.
Nastra Evans, 1955 — 11 spp.
Cymaenes Scudder, 1872 — 27 spp.
Lerema Scudder, 1872 — 8 spp.
Perichares Scudder, 1872 — 9 spp.
Rhinthon Godman in Godman & Salvin, [1900] — 2 spp.
Decinea Evans, 1955 — 10 spp.
Cyclosma Draudt in Seitz, 1923 — 3 spp.
Holguinia Evans, 1955 — 1 sp.
Oeonus Godman in Godman & Salvin, [1900] — 3 spp.
Orthos Evans, 1955 — 5 spp.
Bruna Evans, 1955 — 1 sp.
Conga Evans, 1955 — 5 spp.
Ancyloxypha Felder, 1862 — 7 spp.
Oarisma Scudder, 1872 — 7 spp.
Copaeodes Speyer in Edwards, 1877 — 6 spp.
Adopaeoides Godman in Godman & Salvin, [1900] — 2 spp.

Tribe ?

Adlerodea Hayward, 1940 — 4 spp.
Arita Evans, 1955 — 4 spp.
Cobalopsis Godman in Godman & Salvin, [1900] — 16 spp.
Cumbre Evans, 1955 — 3 spp.
Eogenes Mabille in Seitz, 1909 — 2 spp.
Falga Mabille, 1897 — 4 spp.
Igapophilus Mielke, 1980 — 1 sp.
Inglorius Austin, 1997 — 1 sp.
Lento Evans, 1955 — 19 spp.
Levina Evans, 1955 — 1 sp.
Misius Evans, 1955 — 1 sp.
Mnasilus Godman in Godman & Salvin, [1900] — 1 sp.
Mnasinous Godman in Godman & Salvin, [1900] — 1 sp.
Mnasitheus Godman in Godman & Salvin, [1900] — 11 spp.
Molla Evans, 1955 — 1 sp.
Molo Godman in Godman & Salvin, [1900] — 4 spp.
Sacrator Evans, 1955 — 2 spp.
Styriodes Schaus, 1913 — 8 spp.

Tribe Thymelini

Thymelicus Hübner, [1819] — 11 spp. 1 sp.

Tribe ?

Actinor Watson, 1893 — 1 sp.
Librita Evans, 1955 — 2 spp.
Pseudocopaeodes Skinner & Williams, 1923 — 1 sp.
Quasimellana Burns, 1994 — 24 spp.
Stinga Evans, 1955 — 1 sp.

Tribe Hesperiini

Hesperia Fabricius, 1793 Branded Skippers — 22 spp. 1 sp.
Ochlodes Scudder, 1872 — 21 spp. 1 sp.

Tribe ?

Onespa Steinhauser, 1974 — 1 sp.
Parachoranthus Miller, 1966 — 1 sp.
Polites Scudder, 1872 — 18 spp.
Wallengrenia Berg, 1897 — 3 spp.
Hylephila Billberg, 1820 — 21 spp.
Linka Evans, 1955 — 1 sp.
Pompeius Evans, 1955 — 6 spp.
Atalopedes Scudder, 1872 — 5 spp.
Atrytone Scudder, 1872 — 1 sp.
Anatrytone Dyar, 1905 — 8 spp.
Problema Skinner & Williams, 1924 — 2 spp.

Tribe ?

Buzyges Godman in Godman & Salvin, [1900] — 1 sp.

Tribe Taractrocerini

Taractrocera Butler, [1870] Grass Darts — 14 spp.
Oriens Evans, 1932 Dartlets — 8 spp.
Larva on Imperata cylindrica, Paspalum conjugatum, ;
Potanthus Scudder, 1872 Darts — 36 spp.
Telicota Moore, [1881] Palm Darts — 30 spp.
Larva on Oryza sativa, Saccharum officinarum, Imperata cylindrica, Cocos nucifera, Calamus , ;
Cephrenes Waterhouse & Lyell, 1914 — 6 spp.

Tribe ?

Kobrona Evans, 1935 — 18 spp.
Pastria Evans, 1949 — 3 spp.
Poanes Scudder, 1872 — 16 spp.
Prusiana Evans, 1937 — 3 spp.
Paratrytone Godman in Godman & Salvin, [1900] — 13 spp.
Choranthus Scudder, 1872 — 9 spp.
Halotus Godman in Godman & Salvin, [1900] — 3 spp.
Euphyes Scudder, 1872 — 21 spp.
Arotis Mabille, 1904 — 7 spp.
Chalcone Evans, 1955 — 4 spp.
Hansa Evans, 1955 — 3 spp.
Libra Evans, 1955 — 2 spp.
Metron Godman in Godman & Salvin, [1900] — 6 spp.
Phemiades Hübner, [1819] — 5 spp.
Propertius Evans, 1955 — 2 spp.
Serdis Mabille, 1904 — 3 spp.
Asbolis Mabille, 1904 — 1 sp.
Atrytonopsis Godman in Godman & Salvin, [1900] — 12 spp.
Amblyscirtes Scudder, 1872 — 31 spp.
Lerodea Scudder, 1872 — 10 spp.
Oligoria Scudder, 1872 — 1 sp.
Caligulana Bell, 1942 — 1 sp.
Oxynthes Godman in Godman & Salvin, [1900] — 2 spp.
Calpodes Hübner, [1819] — 1 sp.
Panoquina Hemming, 1934 — 17 spp.
Nyctelius Hayward, 1948 — 2 spp.
Thespieus Godman in Godman & Salvin, [1900] — 32 spp.
Tirynthoides Bell, 1940 — 2 spp.

Tribe Gegenini

Gegenes Hübner, [1819] Pygmy Skippers, Hottentots — 4 spp.
Parnara Moore, [1881] — 10 spp.
Larva on Ozyra sativa, Saccharum officinarum, Zea mays, Bambusa , Colocasia esculenta, ;
Borbo Evans, 1949 Swifts — 21 spp.
Pseudoborbo Lee, 1966 — 1 sp.
Pelopidas Walker, 1870 Millet Skippers — 10 spp.
Polytremis Mabille, 1904 — 19 spp.
Baoris Moore, [1881] Swifts — 8 spp.
Caltoris Swinhoe, 1893 — 19 spp.
Iton de Nicéville, 1895 — 2 spp.

Tribe ?

Brusa Evans, 1937 — 2 spp.
Melphina Evans, 1937 — 11 spp.
Sebastonyma Watson, 1893 — 4 spp.
Sovia Evans, 1949 — 9 spp.
Halpemorpha Huang, Fan & Chiba, 2019 — 4 spp.
Zenonia Evans, 1935 — 3 spp.
Platylesches Holland, 1896 Hoppers — 22 spp.
Fresna Evans, 1937 Acraea Skippers — 6 spp.

Tribe ?

Appia Evans, 1955 — 1 sp.
Kedestes Watson, 1893 Rangers — 24 spp.
Prosopalpus Holland, 1896 — 3 spp.
Gorgyra Holland, 1896 — 22 spp.
Teniorhinus Holland, 1892 — 5 spp.
Xanthodisca Aurivillius, 1925 — 4 spp.
Osmodes Holland, 1892 — 15 spp.
Osphantes Holland, 1896 — 1 sp.
Acada Evans, 1937 — 2 spp.
Ankola Evans, 1937 — 1 sp.
Parosmodes Holland, 1896 — 3 spp.
Rhabdomantis Holland, 1896 — 2 spp.
Acleros Mabille in Grandidier, [1885] — 8 spp.
Semalea Holland, 1896 — 5 spp.
Andronymus Holland, 1896 Darts — 10 spp.
Hypoleucis Mabille, 1891 — 3 spp.
Meza Hemming, 1939 — 10 spp.
Paracleros Berger, 1978 — 6 spp.
Paronymus Aurivillius, 1925 — 5 spp.
Moltena Evans, 1937 — 1 sp.
Monza Evans, 1937 — 3 spp.
Mopala Evans, 1937 — 1 sp.
Perrotia Oberthür, 1916 — 15 spp.
Ploetzia Saalmüller, 1884 — 1 sp.
Pteroteinon Watson, 1893 — 9 spp.
Chondrolepis Mabille, 1904 Snow Horned Skippers — 7 spp.
Gamia Holland, 1896 — 3 spp.
Gretna Evans, 1937 — 8 spp.
Malaza Mabille, 1904 — 3 spp.
Zophopetes Mabille, 1904 Nightfighters — 6 spp.
Artitropa Holland, 1896 Nightfighters — 9 spp.

Group Plastingia

Arnetta Watson, 1893 Bobs — 7 spp.
Suada de Nicéville, 1895 — 4 spp.
Scobura Elwes & Edwards, 1897 — 8 spp.
Suastus Moore, [1881] — 4 spp.
Cupitha Moore, 1884 — 1 sp.
Zographetus Watson, 1893 — 10 spp.
Oerane Elwes & Edwards, 1897 — 1 sp.
Hyarotis Moore, [1881] — 4 spp.
Quedara Swinhoe, 1919 — 6 spp.
Ilma Swinhoe, 1905 — 1 sp.
Isma Distant, 1886 — 18 spp.
Xanthoneura Eliot in Corbet & Pendlebury, 1978 — 3 spp.
Plastingia Butler, 1870 — 7 spp.
Salanoemia Eliot in Corbet & Pendlebury, 1978 — 6 spp.
Pemara Eliot in Corbet & Pendlebury, 1978 — 1 sp.
Pyroneura Eliot in Corbet & Pendlebury, 1978 — 15 spp.
Pseudokerana Eliot in Corbet & Pendlebury, 1978 — 1 sp.
Lotongus Distant, 1886 — 3 spp.
Zela de Nicéville, 1895 — 8 spp.
Gangara Moore, [1881] — 4 spp.
Erionota Mabille, 1878 — 10 spp.
Ge de Nicéville, 1895 — 1 sp.
Matapa Moore, [1881] — 8 spp.
Unkana Distant, 1886 — 4 spp.
Hidari Distant, 1886 — 3 spp.
Eetion de Nicéville, 1895 — 1 sp.
Acerbas de Nicéville, 1895 — 7 spp.
Pirdana Distant, 1886 — 5 spp.

Group ?

Creteus de Nicéville, 1895 — 1 sp.
Fulda Evans, 1937 — 8 spp.
Isoteinon C. & R. Felder, 1862 — 1 sp.
Prada Evans, 1949 — 4 spp.
Praescobura Devyatkin, 2002 — 1 sp.
Pudicitia de Nicéville, 1895 — 1 sp.
Tiacellia Evans, 1949 — 1 sp.

Tribe ?

Baracus Moore, [1881] Hedge Hoppers — 1 sp.

Tribe Ancistroidini

In addition to these, some other genera may belong... -- msa

Astictopterus C. & R. Felder, 1860 Fairies — 9 spp.
Iambrix Watson, 1893 — 2 spp.
Idmon de Nicéville, 1895 — 3 spp.
Koruthaialos Watson, 1893 — 4 spp.
Psolos Staudinger, 1889 — 1 sp.
Stimula de Nicéville, 1898 — 1 sp.
Ancistroides Butler, 1874 — 4 spp.
Notocrypta de Nicéville, 1889 Demons — 13 spp.
Larva on Musa , Curcuma , Costus speciosus, ;
Udaspes Moore, [1881] — 2 spp.

Tribe Ampittiini

Ampittia Moore, [1881] Bush Hoppers — 10 spp.

Tribe Astictopterini

In addition to these, some other genera may belong... -- msa

Aeromachus de Nicéville, 1890 Scrub Hoppers — 20 spp.
Halpe Moore, 1878 Aces — 52 spp.
Pithauria Moore, 1878 — 4 spp.
Onryza Watson, 1893 — 4 spp.
Thoressa Swinhoe, [1913] — 38 spp.

Tribe ?

Aides Billberg, 1820 — 7 spp.
Alera Mabille, 1891 — 6 spp.
Anthoptus Bell, 1942 — 5 spp.
Apaustus Hübner, [1819] — 3 spp.
Argon Evans, 1955 — 1 sp.
Aroma Evans, 1955 — 2 spp.
Arrhenes Evans, 1934 — 7 spp.
Artines Godman in Godman & Salvin, [1901] — 10 spp.
Banta Evans, 1949 — 3 spp.
Caenides Holland, 1896 — 21 spp.
Cantha Evans, 1955 — 7 spp.
Carystina Evans, 1955 — 4 spp.
Carystoides Godman in Godman & Salvin, [1901] — 19 spp.
Carystus Hübner, [1819] — 9 spp.
Ceratrichia Butler, [1870] Forest Sylphs — 19 spp.
Chloeria Mabille, 1904 — 1 sp.
Cobaloides Hayward, 1938 — 1 sp.
Cobalus Hübner, [1819] — 3 spp.
Cravera de Jong, 1983 — 2 spp.
Cynea Evans, 1955 — 16 spp.
Damas Godman in Godman & Salvin, [1901] — 3 spp.
Dion Godman in Godman & Salvin, [1901] — 3 spp.
Dubiella Evans, 1936 — 2 spp.
Ebusus Evans, 1955 — 1 sp.
Enosis Mabille, 1889 — 12 spp.
Eprius Godman in Godman & Salvin, [1901] — 1 sp.
Eutocus Godman in Godman & Salvin, [1901] — 8 spp.
Eutychide Godman in Godman & Salvin, [1900] — 8 spp.
Evansiella Hayward, 1948 — 1 sp.
Flaccilla Godman in Godman & Salvin, [1901] — 1 sp.
Gallio Evans, 1955 — 1 sp.
Joanna Evans, 1955 — 2 spp.
Justinia Evans, 1955 — 8 spp.
Lamponia Evans, 1955 — 3 spp.
Lindra Evans, 1955 — 5 spp.
Lucida Evans, 1955 — 5 spp.
Ludens Evans, 1955 — 3 spp.
Lycas Godman in Godman & Salvin, [1901] — 2 spp.
Lychnuchoides Godman in Godman & Salvin, [1901] — 3 spp.
Lychnuchus Hübner, [1831] — 4 spp.
Megaleas Godman in Godman & Salvin, [1901] — 2 spp.
Methion Godman in Godman & Salvin, [1900] — 1 sp.
Methionopsis Godman in Godman & Salvin, [1901] — 4 spp.
Miltomiges Mabille, 1903 — 1 sp.
Mimene Joicey & Talbot, 1917 — 22 spp.
Mnaseas Godman in Godman & Salvin, [1901] — 2 spp.
Mnasicles Godman in Godman & Salvin, [1901] — 3 spp.
Mnestheus Godman in Godman & Salvin, [1901] — 2 spp.
Moeris Godman in Godman & Salvin, [1900] — 5 spp.
Moeros Evans, 1955 — 1 sp.
Morys Godman in Godman & Salvin, [1900] — 8 spp.
Mucia Godman in Godman & Salvin, [1900] — 4 spp.
Naevolus Hemming, 1939 — 1 sp.
Neochlodes Austin & DeVries, 2001 — 1 sp.
Neoxeniades Hayward, 1938 — 10 spp.
Niconiades Hübner, [1821] — 17 spp.
Nyctus Mabille, 1891 — 2 spp.
Ochus de Nicéville, 1894 — 1 sp.
Ocybadistes Heron, 1894 — 7 spp.
Onophas Godman in Godman & Salvin, [1900] — 3 spp.
Orphe Godman in Godman & Salvin, [1901] — 2 spp.
Orses Godman in Godman & Salvin, [1901] — 2 spp.
Pamba Evans, 1955 — 2 spp.
Panca Evans, 1955 — 1 sp.
Papias Godman in Godman & Salvin, [1900] — 10 spp.
Paracarystus Godman in Godman & Salvin, [1900] — 3 spp.
Pardaleodes Butler, 1870 — 6 spp.
Parphorus Godman in Godman & Salvin, [1900] — 14 spp.
Penicula Evans, 1955 — 7 spp.
Phanes Godman in Godman & Salvin, [1901] — 7 spp.
Pheraeus Godman in Godman & Salvin, [1900] — 11 spp.
Phlebodes Hübner, [1819] — 12 spp.
Propapias Mielke, 1992 — 1 sp.
Pseudosarbia Berg, 1897 — 2 spp.
Psoralis Mabille, 1904 — 11 spp.
Punta Evans, 1955 — 1 sp.
Pyrrhocalles Mabille, 1904 — 2 spp.
Pyrrhopygopsis Godman in Godman & Salvin, [1901] — 4 spp.
Quinta Evans, 1955 — 2 spp.
Racta Evans, 1955 — 5 spp.
Remella Hemming, 1939 — 5 spp.
Repens Evans, 1955 — 1 sp.
Sabera Swinhoe, 1908 — 13 spp.
Saniba Mielke & Casagrande, 2003 — 1 sp.
Saliana Evans, 1955 — 20 spp.
Saturnus Evans, 1955 — 4 spp.
Sodalia Evans, 1955 — 3 spp.
Sucova Evans, 1955 — 1 sp.
Suniana Evans, 1934 — 3 spp.
Jongiana Mielke & Casagrande, 2002 — 1 sp.
Synale Mabille, 1904 — 5 spp.
Talides Hübner, [1819] — 7 spp.
Telles Godman in Godman & Salvin, [1900] — 2 spp.
Tellona Evans, 1955 — 1 sp.
Thargella Godman in Godman & Salvin, [1900] — 2 spp.
Thoon Godman in Godman & Salvin, [1900] — 11 spp.
Thracides Hübner, [1819] — 7 spp.
Tigasis Godman in Godman & Salvin, [1900] — 9 spp.
Tirynthia Godman in Godman & Salvin, [1900] — 3 spp.
Tisias Godman in Godman & Salvin, [1901] — 7 spp.
Tromba Evans, 1955 — 2 spp.
Turesis Godman in Godman & Salvin, [1901] — 6 spp.
Turmada Evans, 1955 — 2 spp.
Vacerra Godman in Godman & Salvin, [1900] — 8 spp.
Vehilius Godman in Godman & Salvin, [1900] — 11 spp.
Venas Evans, 1955 — 2 spp.
Vertica Evans, 1955 — 4 spp.
Vettius Godman in Godman & Salvin, [1901] — 24 spp.
Vinius Godman in Godman & Salvin, [1900] — 5 spp.
Vinpeius Austin, 1997 — 1 sp.
Virga Evans, 1955 — 8 spp.
Wahydra Steinhauser, 1991 — 9 spp.
Yanoancistroides Huang, 1999 — 2 spp.
Xeniades Godman in Godman & Salvin, [1900] — 4 spp.
Zalomes Bell, 1947 — 10 spp.
Zariaspes Godman in Godman & Salvin, [1900] — 2 spp.
Zenis Godman in Godman & Salvin, [1900] — 2 spp.

1000x744 (~95Kb) USA: Mima Mounds Natural Area Preserve, Thurston Co., WA, 28.6.2000, Photo © Markku Savela	1000x774 (~108Kb) USA: Mima Mounds Natural Area Preserve, Thurston Co., WA, 28.6.2000, Photo © Markku Savela	450x358 (~26Kb) Venezuela, Yacambu park near Barquisimeto, July 2002, Photo © Simon Coombes
500x370 (~31Kb) Venezuela, Yacambu park near Barquisimeto, July 2002, Photo © Simon Coombes	500x384 (~38Kb) Venezuela, Merida and the Sierra Nevada National Park, July 2002, Photo © Simon Coombes	400x249 (~22Kb) Venezuela, Parai Tepui near Mount Roraima in the Gran Sabana, July 2002, Photo © Simon Coombes
769x514 (~129Kb) Thailand, Krabi 24.12.2003-7.1.2004, Photo © Olli Vesikko	600x400 (~67Kb) upperside India: Subansiri, 2.11.2005, Photo © Kedar Tokekar	600x400 (~59Kb) underside India: Subansiri, 2.11.2005, Photo © Kedar Tokekar
600x400 (~70Kb) ♂ ♀ underside India: Subansiri, 3.11.2005, Photo © Kedar Tokekar	1040x859 (~271Kb) Ethiopia, Amhara, Bahir Dar, a grassy swamp margin at Tana Hotel. 4th August 2012, Photo © Oleg Kosterin	1025x921 (~300Kb) Ethiopia, Amhara, Bahir Dar, a grassy swamp margin at Tana Hotel. 4th August 2012, Photo © Oleg Kosterin
1296x1326 (~311Kb) upperside Cambodia, Kep Province, Kep, Treetop Buingalow. 6th December 2010, Photo © Oleg Kosterin	872x975 (~201Kb) upperside Cambodia, Kep Province, Kep, Treetop Buingalow. 6th December 2010, Photo © Oleg Kosterin	1182x997 (~239Kb) underside Cambodia, Koh Kong Province, 17 km ENE of Koh Kong, a forest rivulet, 23rd May 2013, Photo © Oleg Kosterin

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.

Unknown or unplaced taxa

Prenes devantieri Mabille, 1904; (nom. nud.); TL: Brazil(Rio Grande do Sul)

devantieri; [NL4A], 85

Pamphila mago Ménétriés, 1857; (nom. nud.); TL: Venezuela

mago; [NL4A], 85

Calpodes orthamas Weeks, 1901; (nom. nud.); TL: Bolivia

orthamas; [NL4A], 85

Atrytone pericia Sheldon, 1938; (nom. nud.); TL: Tobago

pericia; [NL4A], 85

Hesperia powelli Stanford & Opler, 1993; (nom. nud.); TL: Mexico(Chihuahua)

powelli; [NL4A], 85

Pamphila pyrgina Capronnier, 1881; (nom. nud.); TL: Brazil(Rio de Janeiro)

pyrgina; [NL4A], 85

Prenes quonga Hall, 1939; (nom. nud.); TL: Guyana

quonga; [NL4A], 85

Megistias semiargus J. & W. Zikán, 1968; (nom. nud.); TL: Brazil(Rio de Janeiro/Minas Gerais)

semiargus; [NL4A], 85

Pamphila valva Weeks, 1901; (nom. nud.); TL: Bolivia

valva; [NL4A], 85

12.9.2023 (2034)

References:

[CoL] Catalogue of Life
Bánki, O., Roskov, Y., Döring, M., Ower, G., Hernández Robles, D. R., Plata Corredor, C. A., Stjernegaard Jeppesen, T., Örn, A., Vandepitte, L., Hobern, D., Schalk, P., DeWalt, R. E., Ma, K., Miller, J., Orrell, T., Aalbu, R., Abbott, J., Adlard, R., Aedo, C., et al. (2024). Catalogue of Life Checklist (Version 2024-03-26); https://doi.org/10.48580/dfz8d
[EVERSMANNIA]
Eversmannia contents
[Kedar Tokekar]
India
[Oleg Kosterin] Institue of Cytology & Genetics
Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Olli Vesikko]
http://koti.welho.com/ovesikko/
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[iNaturalist]
iNaturalist

Some related literature:

Korb & Bolshakov, 2011
[A catalogue of butterflies (Lepidoptera: Papilioformes) of the former USSR. Second edition, reformatted and updated] (in Russian) Eversmannia Suppl. 2 : 1-123
[NL4A]; Lamas, 2004
Checklist: Part 4A, Hesperioidea - Papilionoidea Atlas of Neotropical Lepidoptera
[BK]; Larsen, 1996
The Butterflies of Kenya and their natural history Butterflies of Kenya
[RFEL]; Leley, 2016
Annotated catalogue of the insects of Russian Far East. Volume II. Lepidoptera Cat. ins. Russian Far East. 2 : 1-812

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.