Life Insecta Lepidoptera Ditrysia Papilionoidea Nymphalidae Nymphalinae

700x579 (~85Kb) USA: Bridge Creek Trailhead, North Cascades Highway, (48°30'30N 120°43'00W), Chelan Co., WA, 28.7.1999, Photo © Markku Savela	1000x720 (~149Kb) USA: Near Crystal Mtn, (46°57'51N 121°28'55W), Pierce Co., WA, 27.7.1999, Photo © Markku Savela	600x639 (~46Kb) USA: Wenatchee National Forest, (47°04'12N 120°56'10W), Kittitas Co., WA, 29.7.1999, Photo © Markku Savela
900x663 (~71Kb) USA: Wenatchee National Forest, (47°04'12N 120°56'10W), Kittitas Co., WA, 29.7.1999, Photo © Markku Savela	900x588 (~60Kb) USA: Wenatchee National Forest, (47°04'12N 120°56'10W), Kittitas Co., WA, 29.7.1999, Photo © Markku Savela	700x919 (~101Kb) ♂ ♀ USA: Corral Pass region (47°00'15N 121°29'02W), Pierce Co., WA, 18.7.2000, Photo © Markku Savela
660x784 (~82Kb) ♂ ♀ USA: Corral Pass region (47°00'15N 121°29'02W), Pierce Co., WA, 18.7.2000, Photo © Markku Savela	1800x1539 (~254Kb) ♂ upperside USA: Washington, Kittitas Co., LepSoc 2010 Field Trip to Red Top Mtn (meeting point, on FR 9738?), 8.7.2010, Photo © Markku Savela

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.

• Euphydryas phaeton (Drury, [1773]) ENA. See [maps] Baltimore

  • Papilio phaeton Drury, [1773]; Illust. Nat. Hist. Exot. Insects 1: index, 42, pl. 21, f. 3-4; TL: New York
  • phaedon Herrich-Schäffer, 1865; (misspel.)

  • Euphydryas (Euphydryas) phaeton phaeton; [P&N23]
  • Euphydryas (Euphydryas) phaeton; Pelham, 2008, J. Res. Lepid. 40: 362; [P&N23]
  • Euphydryas phaeton; Dyar, 1903, Bull. U.S. nat. Mus. 52: 17; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [NACL], #4516; [NAB], 602; [EBW]; [BOW]: pl. 13, f. 28; [Opler]; [NL4A], #2064; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA] ; [BAMONA] ; [GALEPS]

  432x289 (~30Kb) USA: Ann Arbor, Michigan, May 1999, Photo © Christopher A. Rickards	1036x991 (~183Kb) USA: Alabama, Jeff. Co., 13.5.2000, Photo © Vitaly Charny	1157x771 (~131Kb) larva USA: Alabama, 22.4.2001, Photo © Vitaly Charny
  1091x867 (~104Kb) larva USA: Alabama, 5.5.2001, Photo © Vitaly Charny	800x1238 (~239Kb) larva USA: Alabama, 5.5.2001, Photo © Vitaly Charny	1300x927 (~250Kb) upperside USA: Alabama, 5.5.2001, Photo © Vitaly Charny
  967x1339 (~220Kb) underside USA: Alabama, 5.5.2001, Photo © Vitaly Charny	828x633 (~95Kb) USA: Alabama, 13.5.2001, Photo © Vitaly Charny	1049x766 (~209Kb) upperside USA: Alabama, 6.5.2006, Photo © Vitaly Charny

  Larva on Chelone glabra, Gerardia grandiflora, G. pedicularia, Plantago lanceolata, Fraxinus americana [NAB]

  • E. p. phaeton New York, Maryland, Maine, ...

    • Melitaea phaëtaena Hübner, [1819]; Verz. bek. Schmett. (2): 28; TL: New York
      • = ; [NL4A], #2064a; Pelham, 2008, J. Res. Lepid. 40: 362
    • Argynnis phaetontea Godart, 1819; Encyclopédie Méthodique 9 (1): 288; TL: New York
      • = ; [NL4A], #2064a
    • Melitaea phaeton ab. ♂ superba Strecker, 1878; Synon. Cat. Macroplep.: 125; TL: New York
      • = ; [NL4A], #2064a; Pelham, 2008, J. Res. Lepid. 40: 362
    • Melitaea phaeton ab. phaethusa Hulst, 1881; Bull. Brooklyn Ent. Soc. 3 (9): 77; TL: New York
      • = ; [NL4A], #2064a
      • = (phaetusa); Pelham, 2008, J. Res. Lepid. 40: 362
    • Melitaea phaeton ab. streckeri Ellsworth, 1902; Ent. News. 13: 104; TL: New York
      • = ; [NL4A], #2064a
      • = (phaetusa); Pelham, 2008, J. Res. Lepid. 40: 362
    • Euphydryas phaëton schausi Clark, 1927; Proc. U.S. Nat. Mus. 71 (2683): 1, pl. 1; TL: Maryland
      • = ; [NL4A], #2064a; Pelham, 2008, J. Res. Lepid. 40: 362
    • Euphydryas phaëton ab. magnifica Clark, 1927; Proc. U.S. Nat. Mus. 71 (2683): 7, pl. 2, f. 9-12; TL: Maryland
      • = ; [NL4A], #2064a; Pelham, 2008, J. Res. Lepid. 40: 363
    • Euphydryas phaeton borealis Chermock & Chermock, 1940; Proc. Penn. Acad. Sci. 14: 140; TL: Lincoln, Maine
      • = ; [NL4A], #2064a; Pelham, 2008, J. Res. Lepid. 40: 362

    • Euphydryas (Euphydryas) phaeton phaeton; Pelham, 2008, J. Res. Lepid. 40: 362
    • Euphydryas phaeton borealis; [LepIndex]; [CoL]
    • Euphydryas phaeton phaeton; [NL4A], #2064a; [iNaturalist] ; [BOA]
    • Euphydryas phaeton streckeri; [LepIndex]; [CoL]
    • Euphydryas phaeton superba; [LepIndex]; [CoL]
  • E. p. ozarkae Masters, 1968

    • Euphydryas phaeton ozarkae Masters, 1968; Ent. News 79 (4): 87

    • Euphydryas (Euphydryas) phaeton ozarkae; Pelham, 2008, J. Res. Lepid. 40: 363; [P&N23]
    • Euphydryas phaeton ozarkae; [NACL], #4516a; [LepIndex]; [CoL] ; [iNaturalist] ; [BOA]
• Euphydryas anicia (Doubleday, [1847]) British Columbia. See [maps] Anicia Checkerspot

  • Melitaea anicia Doubleday, [1847]; Gen. diurn. Lep. (1): 179

  • Euphydryas (Euphydryas) anicia anicia; [P&N23]
  • Euphydryas (Euphydryas) anicia; Pelham, 2008, J. Res. Lepid. 40: 367; [P&N23]
  • Euphydryas anicia; [NAB], 606; [BOW]: pl. 13, f. 23; [Opler]; [NL4A], #2061; [MPG] ; [iNaturalist] ; [LEPIFORUM] ; [BOA] ; [BOA] ; [BOA] ; [BAMONA]
  • Euphydryas chalcedona anicia;
  • Lemonias anicia; Dyar, 1903, Bull. U.S. nat. Mus. 52: 17
  • Occidryas anicia; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [NACL], #4519; [LepIndex] ; [CoL]

  identification uncertain 1300x1013 (~240Kb) USA: Cabin Mountain, (1408m), (47°14'15N 121°16'27W), Kittitas Co., WA, 29.7.2006, Photo © Markku Savela

  identification uncertain 1200x1685 (~234Kb) USA: Cabin Mountain, (1408m), (47°14'15N 121°16'27W), Kittitas Co., WA, 29.7.2006, Photo © Markku Savela

  Larva on Castilleja , Penstemon , Scrophulariaceae , Plantaginaceae , Boraginaceae [NAB]

  • E. a. anicia

    • Euphydryas anicia tr.f. mayi Gunder, 1932; Pan-Pacific Ent. 8 (3): 127
      • = ; Pelham, 2008, J. Res. Lepid. 40: 367

    • Euphydryas (Euphydryas) anicia anicia; Pelham, 2008, J. Res. Lepid. 40: 367
    • Euphydryas anicia anicia; [iNaturalist] ; [BOA]
  • E. a. helvia (Scudder, 1869) Alaska

    • Melitaea helvia Scudder, 1869; Proc. Boston Soc. nat. Hist. 12: 405

    • Euphydryas (Euphydryas) anicia helvia; Pelham, 2008, J. Res. Lepid. 40: 367; [P&N23]
    • Euphydryas anicia helvia; [iNaturalist] ; [BOA]
    • Lemonias helvia; Dyar, 1903, Bull. U.S. nat. Mus. 52: 18
    • Occidryas anicia helvia; [NACL], #4519a; [LepIndex]; [CoL]
  • E. a. eurytion (Mead, 1875) Colorado

    • Melitaea eurytion Mead, 1875; in Wheeler, Reports upon Zool. coll. (8): 759 (Edw., MS)
    • Melitaea anicia capella ab. rubrolimbata Comstock, 1918; Lepidopterist 2 (5): 36; TL: South Park, Colorado
      • = ; Pelham, 2008, J. Res. Lepid. 40: 368
    • Euphydryas anicia carolae Emmel & Harris, 1998; Syst. W. N. Am. Butts. (32): 433
      • = ; Pelham, 2008, J. Res. Lepid. 40: 368

    • Euphydryas (Euphydryas) anicia eurytion; Pelham, 2008, J. Res. Lepid. 40: 368; [P&N23]
    • Euphydryas anicia eurytion; [iNaturalist] ; [BOA]
    • Occidryas anicia eurytion; [NACL], #4519i; [LepIndex]; [CoL]
  • E. a. wheeleri (H. Edwards, 1881) S.Nevada

    • Melitaea anicia var. wheeleri H. Edwards, 1881; Papilio 1 (4): 52

    • Euphydryas (Euphydryas) anicia wheeleri; Pelham, 2008, J. Res. Lepid. 40: 379; [P&N23]
    • Euphydryas anicia wheeleri; [iNaturalist] ; [BOA]
    • Lemonias nubigena wheeleri; Dyar, 1903, Bull. U.S. nat. Mus. 52: 17
    • Occidryas anicia wheeleri; [NACL], #4519p; [LepIndex]; [CoL]
  • E. a. brucei (Edwards, 1888)

    • Melitaea brucei Edwards, 1888; Can. Ent. 20 (5): 81
    • Melitaea anicia anica ab. melanodisca Comstock, 1918; Lepidopterist 2 (5): 34, pl. 6, f. 1; TL: Hall Valley, Colorado
      • = ; Pelham, 2008, J. Res. Lepid. 40: 368

    • Euphydryas (Euphydryas) anicia brucei; Pelham, 2008, J. Res. Lepid. 40: 368; [P&N23]
    • Euphydryas anicia brucei; [iNaturalist] ; [BOA]
    • Lemonias anicia brucei; Dyar, 1903, Bull. U.S. nat. Mus. 52: 17
  • E. a. capella (Barnes, 1897) Colorado

    • Melitaea nubigena var. capella Barnes, 1897; Can. Ent. 29 (2): 41
    • Euphydryas anicia var. capella ab. ♂ oslari Gunder, 1925; Ent. News 36 (7): 196, pl. 5, f. 7; TL: Platte Canyon, Colorado
      • = ; Pelham, 2008, J. Res. Lepid. 40: 369

    • Euphydryas (Euphydryas) anicia capella; Pelham, 2008, J. Res. Lepid. 40: 369; [P&N23]
    • Euphydryas anicia capella; [iNaturalist] ; [BOA]
    • Occidryas anicia capella; [NACL], #4519c; [LepIndex]; [CoL]
  • E. a. maria (Skinner, 1899) Utah

    • Melitaea maria Skinner, 1899; Can. Ent. 31 (1): 12; TL: Park City, Utah
    • Euphydryas maria tr.f. skinneri Gunder, 1928; Can. Ent. 60 (7): 164, pl. A, f. 9; TL: Eureka, Utah
      • = ; Pelham, 2008, J. Res. Lepid. 40: 368
    • Euphydryas maria tr.f. spaldingi Gunder, 1928; Can. Ent. 60 (7): 165, pl. A, f. 10; TL: Stockton, Utah
      • = ; Pelham, 2008, J. Res. Lepid. 40: 368

    • Euphydryas (Euphydryas) anicia maria; Pelham, 2008, J. Res. Lepid. 40: 367; [P&N23]
    • Euphydryas anicia maria; [iNaturalist] ; [BOA]
    • Lemonias maria; Dyar, 1903, Bull. U.S. nat. Mus. 52: 17
    • Occidryas anicia maria; [NACL], #4519j; [LepIndex]; [CoL]
  • E. a. hermosa (Wright, 1905) Arizona

    • Melitaea hermosa Wright, 1905; Butts. West Coast US: 157, pl. 19, f. 179; TL: Arizona
    • Euphydryas anicia wheeleri tr.f. duncani Gunder, 1934; Can. Ent. 66 (6): 129; TL: Santa Catalina Mts, Arizona
      • = ; Pelham, 2008, J. Res. Lepid. 40: 368

    • Euphydryas (Euphydryas) anicia hermosa; Pelham, 2008, J. Res. Lepid. 40: 368; [P&N23]
    • Euphydryas anicia hermosa; [NL4A], #2061a; [iNaturalist] ; [BOA]
    • Occidryas anicia hermosa; [NACL], #4519d; [LepIndex]; [CoL]
  • E. a. magdalena Barnes & McDunnough, 1918

    • Euphydryas magdalena Barnes & McDunnough, 1918; Contr. nat. Hist. Lepid. N. Am. 4 (2): 71, pl. 11, f. 1-4; TL: White Mtns, Arizona

    • Euphydryas (Euphydryas) anicia magdalena; Pelham, 2008, J. Res. Lepid. 40: 368; [P&N23]
    • Euphydryas anicia magdalena; [iNaturalist] ; [BOA]
    • Occidryas anicia magdalena; [NACL], #4519e; [LepIndex]; [CoL]
  • E. a. bernadetta Leussler, 1920 Nebraska

    • Euphydryas bernadetta Leussler, 1920; Ent. News 31 (4): 102; TL: Monroe Canyon, nr Harrison, Sioux Co., Nebraska
    • Euphydryas bernadetta ♂ tr.f. leussleri Gunder, 1927; Ent. News. 38 (5): 136, pl. 2, f. 8; TL: nr Harrison, Sioux Co., Nebraska
      • = ; Pelham, 2008, J. Res. Lepid. 40: 369
    • Euphydryas anicia r. bernadetta tr.f. belli Gunder, 1929; Pan-Pacific Ent. 6 (1): 4; TL: Harrison, Sioux Co., Nebraska
      • = ; Pelham, 2008, J. Res. Lepid. 40: 369

    • Euphydryas (Euphydryas) anicia bernadetta; Pelham, 2008, J. Res. Lepid. 40: 369; [P&N23]
    • Euphydryas anicia bernadetta; [iNaturalist] ; [BOA]
    • Occidryas anicia bernadetta; [NACL], #4519m; [LepIndex]; [CoL]
  • E. a. alena Barnes & Benjamin, 1926 Utah

    • Euphydryas alena Barnes & Benjamin, 1926; Bull. South. Calif. Acad. Sci. 25 (3): 93; TL: So. Utah

    • Euphydryas (Euphydryas) anicia alena; Pelham, 2008, J. Res. Lepid. 40: 368; [P&N23]
    • Euphydryas anicia alena; [iNaturalist] ; [BOA]
    • Occidryas anicia alena; [NACL], #4519f; [LepIndex]; [CoL]
  • E. a. carmentis Barnes & Benjamin, 1926 Colorado

    • Euphydryas carmentis Barnes & Benjamin, 1926; Bull. South. Calif. Acad. Sci. 25 (3): 93; TL: Pagosa Springs, Colorado
    • Euphydryas carmentis tr.f. charlottae Gunder, 1928; Can. Ent. 60 (7): 165, pl. B, f. 11; TL: Pagosa Springs, Archuleta Co., Colorado
      • = ; Pelham, 2008, J. Res. Lepid. 40: 369

    • Euphydryas (Euphydryas) anicia carmentis; Pelham, 2008, J. Res. Lepid. 40: 369; [P&N23]
    • Euphydryas anicia carmentis; [iNaturalist] ; [BOA]
    • Occidryas anicia carmentis; [NACL], #4519g; [LepIndex]; [CoL]
  • E. a. morandi Gunder, 1928 Nevada

    • Euphydryas morandi Gunder, 1928; Ent. News 39 (10): 304, pl. 14; TL: Kyle Canyon, Mt. Charleston Peak in Spring Mntn, Clark Co., Nevada

    • Euphydryas (Euphydryas) anicia morandi; Pelham, 2008, J. Res. Lepid. 40: 370; [P&N23]
    • Euphydryas anicia morandi; [iNaturalist] ; [BOA]
    • Occidryas anicia morandi; [NACL], #4519r; [LepIndex]; [CoL]
  • E. a. windi Gunder, 1932 Wyoming

    • Euphydryas anicia r. windi Gunder, 1932; Can. Ent. 64 (12): 283; TL: Timber Is., Teton Mts, Teton Co., Wyoming

    • Euphydryas (Euphydryas) anicia windi; Pelham, 2008, J. Res. Lepid. 40: 367; [P&N23]
    • Euphydryas anicia windi; [iNaturalist] ; [BOA]
    • Occidryas anicia windi; [NACL], #4519h; [LepIndex]; [CoL]
  • E. a. hopfingeri Gunder, 1934 N.Washington

    • Euphydryas anicia r. hopfingeri Gunder, 1934; Can. Ent. 66 (6): 128; TL: Black Canyon, Methow Valley, Okanogan Co., Washington
    • Euphydryas anicia hopfingeri tr.f. andersoni Gunder, 1934; Can. Ent. 66 (6): 128; TL: Black Canyon, nr Brewster, Okanogan Co., Washington
      • = ; Pelham, 2008, J. Res. Lepid. 40: 367

    • Euphydryas (Euphydryas) anicia hopfingeri; Pelham, 2008, J. Res. Lepid. 40: 367; [P&N23]
    • Euphydryas anicia hopfingeri; [iNaturalist] ; [BOA]
    • Occidryas anicia hopfingeri; [NACL], #4519k; [LepIndex]; [CoL]
  • E. a. bakeri Stallings & Turner, 1945 Oregon

    • Euphydryas anicia bakeri Stallings & Turner, 1945; J. Kansas Ent. Soc. 18 (2): 82; TL: Cave Creek, Durkee, Oregon

    • Euphydryas (Euphydryas) anicia bakeri; Pelham, 2008, J. Res. Lepid. 40: 367; [P&N23]
    • Euphydryas anicia bakeri; [iNaturalist] ; [BOA]
    • Occidryas anicia bakeri; [NACL], #4519o; [LepIndex]; [CoL]
  • E. a. effi Stallings & Turner, 1945 Idaho

    • Euphydryas anicia effi Stallings & Turner, 1945; J. Kansas Ent. Soc. 18 (2): 82; TL: Corral Creek, Ketchum, Idaho

    • Euphydryas (Euphydryas) anicia effi; Pelham, 2008, J. Res. Lepid. 40: 367; [P&N23]
    • Euphydryas anicia effi; [BOA]
    • Occidryas anicia effi; [NACL], #4519k; [LepIndex]; [CoL]
  • E. a. howlandi Stallings & Turner, 1947 Montana

    • Euphydryas anicia howlandi Stallings & Turner, 1947; Can. Ent. 78 (7-8): 137; TL: Polaris, Montana

    • Euphydryas (Euphydryas) anicia howlandi; Pelham, 2008, J. Res. Lepid. 40: 367; [P&N23]
    • Euphydryas anicia howlandi; [BOA]
    • Occidryas anicia howlandi; [NACL], #4519b; [LepIndex]; [CoL]
  • E. a. macyi Fender & Jewett, 1953 Oregon

    • Euphydryas anicia macyi Fender & Jewett, 1953; Wasmann J. Biol. 11 (1): 115; TL: Wildhorse Creek, Alvord Basin, Harney Co., Oregon

    • Euphydryas (Euphydryas) anicia macyi; Pelham, 2008, J. Res. Lepid. 40: 370; [P&N23]
    • Euphydryas anicia macyi; [iNaturalist] ; [BOA]
    • Occidryas anicia macyi; [NACL], #4519p; [LepIndex]; [CoL]
  • E. a. veazieae Fender & Jewett, 1953 Oregon

    • Euphydryas anicia veazieae Fender & Jewett, 1953; Wasmann J. Biol. 11 (1): 117; TL: Jackass Mtns, Harney Co., Oregon

    • Euphydryas (Euphydryas) anicia veazieae; Pelham, 2008, J. Res. Lepid. 40: 369; [P&N23]
    • Euphydryas anicia veazieae; [iNaturalist] ; [BOA]
    • Occidryas anicia veazieae; [NACL], #4519n; [LepIndex]; [CoL]
  • E. a. chuskae (Ferris & Holland, 1980) New Mexico

    • Occidryas anicia chuskae Ferris & Holland, 1980; Bull. Allyn Mus. 57: 2

    • Euphydryas (Euphydryas) anicia chuskae; Pelham, 2008, J. Res. Lepid. 40: 369; [P&N23]
    • Euphydryas anicia chuskae; [iNaturalist] ; [BOA]
    • Occidryas anicia chuskae; [LepIndex]; [CoL]
  • E. a. cloudcrofti (Ferris & Holland, 1980) New Mexico

    • Occidryas anicia cloudcrofti Ferris & Holland, 1980; Bull. Allyn Mus. 57: 3

    • Euphydryas (Euphydryas) anicia cloudcrofti; Pelham, 2008, J. Res. Lepid. 40: 369; [P&N23]
    • Euphydryas anicia cloudcrofti; [iNaturalist] ; [BOA]
    • Occidryas anicia cloudcrofti; [LepIndex]; [CoL]
  • E. a. variicolor Baughman & Murphy, 1998 California

    • Euphydryas chalcedona variicolor Baughman & Murphy, 1998; Sys. W. N. Am. Butts. (29): 400; TL: California, Mono Co.

    • Euphydryas (Euphydryas) anicia variicolor; Pelham, 2008, J. Res. Lepid. 40: 370; [P&N23]
    • Euphydryas anicia variicolor; [iNaturalist] ; [BOA]
  • E. a. wecoeut Fisher, Spomer & Scott, 2006 Colorado

    • Euphydryas anicia wecoeut Fisher, Spomer & Scott, 2006; Papilio (n.s.) 12: 31, pl. 5; TL: Coal Mine Point, Black Ridge, 7500', Mesa Co., Colorado

    • Euphydryas (Euphydryas) anicia wecoeut; Pelham, 2008, J. Res. Lepid. 40: 368; [P&N23]
    • Euphydryas anicia wecoeut; [iNaturalist] ; [BOA]
• Euphydryas chalcedona (Doubleday, [1847]) W.USA. See [maps] Chalcedon Checkerspot

  • Melitaea chalcedona Doubleday, [1847]; Gen. diurn. Lep. (1): 180; TL: "Haiti"

  • Euphydryas (Euphydryas) chalcedona chalcedona; [P&N23]
  • Euphydryas (Euphydryas) chalcedona; Pelham, 2008, J. Res. Lepid. 40: 363; [P&N23]
  • Euphydryas chalcedon[sic]; [BOW]: pl. 13, f. 25 (missp.)
  • Euphydryas chalcedona; [NAB], 603; [EBW]; [Wahlberg]; [Opler]; [NL4A], #2062; [MPG] ; [iNaturalist] ; [BOA] ; [BAMONA]
  • Lemonias chalcedon[sic]; Dyar, 1903, Bull. U.S. nat. Mus. 52: 17
  • Occidryas chalcedona; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [NACL], #4517; [LepIndex] ; [CoL]

  Larva on Plantago , Lonicera , Symphoricarpos ablus [NAB]

  • E. c. chalcedona California

    • Melitaea cooperi Behr, 1863; Proc. Calif. Acad. Sci. 3 (1): 90; TL: California
      • = ; [NL4A], #2062a; Pelham, 2008, J. Res. Lepid. 40: 364
    • Melitaea chalcedon[sic] ab. fusimacula Barnes, 1900; Can. Ent. 32 (2): 42; TL: California
      • = ; [NL4A], #2062a; Pelham, 2008, J. Res. Lepid. 40: 364
    • Melitaea chalcedon[sic] ab. mariana Barnes, 1900; Can. Ent. 32 (2): 42; TL: California
      • = ; [NL4A], #2062a; Pelham, 2008, J. Res. Lepid. 40: 364
    • Lemonias chalcedon[sic] ab. grundeli Coolidge, 1908; Can. Ent. 40 (11): 42; TL: California
      • = ; [NL4A], #2062a; Pelham, 2008, J. Res. Lepid. 40: 365
    • Melitaea chalcedon[sic] ab. lorquini Oberthür, 1914; Étud. Lépid. Comp. 9 (2): 76, pl. 259, f. 2170; TL: California
      • = ; [NL4A], #2062a; Pelham, 2008, J. Res. Lepid. 40: 365
    • Euphydryas chalcedona ab. omniluteofuscus Gunder, 1925; Ent. News 36 (7): 195, pl. 5, f. 4; TL: California
      • = ; [NL4A], #2062a; Pelham, 2008, J. Res. Lepid. 40: 365
    • Euphydryas chalcedona ab. ♂ hemiluteofuscus Gunder, 1925; Ent. News 36 (7): 196, pl. 5, f. 5; TL: California
      • = ; [NL4A], #2062a; Pelham, 2008, J. Res. Lepid. 40: 365
    • Euphydryas chalcedona ab. supranigrella Comstock, 1926; Bull. South. Calif. Acad. Sci. 25 (1): 30; TL: Mt Wilson, California
      • = ; [NL4A], #2062a; Pelham, 2008, J. Res. Lepid. 40: 365
    • Euphydryas chalcedona ab. suprafusa Comstock, 1926; Bull. South. Calif. Acad. Sci. 25 (1): 30; TL: Los Angeles Co., California
      • = ; [NL4A], #2062a; Pelham, 2008, J. Res. Lepid. 40: 365
    • Euphydryas chalcedona ab. hemimelanica Comstock, 1926; Bull. South. Calif. Acad. Sci. 25 (1): 31; TL: Sulphur Mountain Springs, Ventura Co., California
      • = ; [NL4A], #2062a; Pelham, 2008, J. Res. Lepid. 40: 365
    • Euphydryas chalcedona ab. fusisecunda Comstock, 1926; Bull. South. Calif. Acad. Sci. 25 (1): 31; TL: Mt Wilson, California
      • = ; [NL4A], #2062a; Pelham, 2008, J. Res. Lepid. 40: 365

    • Euphydryas (Euphydryas) chalcedona chalcedona; Pelham, 2008, J. Res. Lepid. 40: 364
    • Euphydryas chalcedona chalcedona; [NL4A], #2062a; [iNaturalist] ; [BOA]
    • Lemonias chalcedon[sic] fusimacula; Dyar, 1903, Bull. U.S. nat. Mus. 52: 17
    • Lemonias chalcedon[sic] mariana; Dyar, 1903, Bull. U.S. nat. Mus. 52: 17
    • Lemonias cooperi; Dyar, 1903, Bull. U.S. nat. Mus. 52: 17
    • Occidryas chalcedona cooperi; [LepIndex]; [CoL]
  • E. c. dwinellei (H. Edwards, 1881)

    • Melitaea chalcedon[sic] var. dwinellei H. Edwards, 1881; Papilio 1 (4): 51

    • Euphydryas (Euphydryas) chalcedona dwinellei; Pelham, 2008, J. Res. Lepid. 40: 364; [P&N23]
    • Euphydryas chalcedona dwinellei; [iNaturalist] ; [BOA]
    • Lemonias chalcedon[sic] dwinellei; Dyar, 1903, Bull. U.S. nat. Mus. 52: 17
    • Occidryas chalcedona dwinnellei[sic, dwinellei]; [NACL], #4517a
  • E. c. macglashanii (Rivers, 1888) California

    • Melitaea macglashanii Rivers, 1888; Proc. Cal. Acad. Sci. (2) 1: 103
    • Euphydryas chalcedona r. mcglashanii f. truckeensis Gunder, 1928; Ent. News 39 (10): 305; TL: Truckee, Calif.
      • = ; Pelham, 2008, J. Res. Lepid. 40: 364
    • Euphydryas chalcedona r. mcglashanii f.tr. hilli Gunder, 1928; Ent. News 39 (10): 306, pl. 14, f. C, D; TL: Auburn, California
      • = ; Pelham, 2008, J. Res. Lepid. 40: 364

    • Euphydryas (Euphydryas) chalcedona macglashanii; Pelham, 2008, J. Res. Lepid. 40: 364; [P&N23]
    • Euphydryas chalcedona macglashanii; [iNaturalist] ; [BOA]
    • Euphydryas macglashanii; [BOW]: pl. 13, f. 26
    • Limonias chalcedon macglashani; Dyar, 1903, Bull. U.S. nat. Mus. 52: 17
    • Occidryas chalcedona macglashanii; [NACL], #4517f; [LepIndex]; [CoL]
    • Occidryas mcglashanii[sic, macglashanii]; McDunnough, 1938
  • E. c. olancha (Wright, 1905)

    • Melitaea olancha Wright, 1905; Butts. West Coast US: 160, pl. 18, f. 160
    • Euphydryas chalcedona r. olancha ♀ tr.f. malcolmi Gunder, 1927; Ent. News. 38 (5): 134, pl. 2, f. 3
      • = ; Pelham, 2008, J. Res. Lepid. 40: 364
    • Euphydryas chalcedona olanca f. georgei Gunder, 1928; Ent. News 39 (10): 306
      • = ; Pelham, 2008, J. Res. Lepid. 40: 364

    • Euphydryas (Euphydryas) chalcedona olancha; Pelham, 2008, J. Res. Lepid. 40: 364; [P&N23]
    • Euphydryas chalcedona olancha; [iNaturalist] ; [BOA]
    • Melitaea clancha; [LepIndex] ; [CoL]
    • Occidryas chalcedona olancha; [NACL], #4517g; [LepIndex]; [CoL]
  • E. c. sierra Wright, 1905

    • Melitaea sierra Wright, 1905; Butts. West Coast US: 151, pl. 18, f. 163
    • Euphydryas sierra ab. ♂ magdalenae Gunder, 1925; Ent. News 36 (7): 197, pl. 5, f. 8; TL: Glen Alpine Creek, El Dorao Co., California
      • = ; Pelham, 2008, J. Res. Lepid. 40: 363
    • Euphydryas sierra ab. umbrobasana Comstock, 1926; Bull. South. Calif. Acad. Sci. 25 (1): 31
      • = ; Pelham, 2008, J. Res. Lepid. 40: 363
    • Euphydryas irelandi Gunder, 1929; Can. Ent. 61 (2): 44
      • = ; Pelham, 2008, J. Res. Lepid. 40: 363

    • Euphydryas (Euphydryas) chalcedona sierra; Pelham, 2008, J. Res. Lepid. 40: 363; [P&N23]
    • Euphydryas chalcedona sierra; [iNaturalist] ; [BOA]
    • Occidryas chalcedona irelandi; [LepIndex]; [CoL]
    • Occidryas chalcedona sierra; [NACL], #4517h; [LepIndex]; [CoL]
  • E. c. klotsi dos Passos, 1938 Arizona

    • Euphydryas chalcedona klotsi dos Passos, 1938; Can. Ent. 70 (10): 199; TL: Roosevelt Lake, Arizona
    • Euphydryas anicia r. hermosa tr.f. venusta Gunder, 1932; Pan-Pacific Ent. 8 (3): 127
      • = ; Pelham, 2008, J. Res. Lepid. 40: 363

    • Euphydryas (Euphydryas) chalcedona klotsi; Pelham, 2008, J. Res. Lepid. 40: 363; [P&N23]
    • Euphydryas chalcedona klotsi; [iNaturalist] ; [BOA]
    • Occidryas chalcedona klotsi; [NACL], #4517c; [LepIndex]; [CoL]
  • E. c. corralensis Emmel & Emmel, 1973

    • Euphydryas chalcedona corralensis Emmel & Emmel, 1973; J. Res. Lepid. 11 (3): 143

    • Euphydryas (Euphydryas) chalcedona corralensis; Pelham, 2008, J. Res. Lepid. 40: 363; [P&N23]
    • Euphydryas chalcedona corralensis; [iNaturalist] ; [BOA]
    • Occidryas chalcedona corralensis; [NACL], #4517d; [LepIndex]; [CoL]
  • E. c. hennei Scott, 1981 California

    • Euphydryas chalcedona hennei Scott, 1981; Papilio (n.s.) 1: 4; TL: California
    • Euphydryas chalcedona quino tr.f. hennei Gunder, 1932; Pan-Pacific Ent. 8 (3): 126; TL: California
      • = ; [NL4A], #2062b; Pelham, 2008, J. Res. Lepid. 40: 363

    • Euphydryas (Euphydryas) chalcedona hennei; Pelham, 2008, J. Res. Lepid. 40: 363; [P&N23]
    • Euphydryas chalcedona hennei; [NL4A], #2062b; [iNaturalist] ; [BOA]
  • E. c. kingstonensis Emmel & Emmel, 1973

    • Euphydryas chalcedona kingstonensis Emmel & Emmel, 1973; J. Res. Lepid. 11 (3): 141

    • Euphydryas (Euphydryas) chalcedona kingstonensis; Pelham, 2008, J. Res. Lepid. 40: 363; [P&N23]
    • Euphydryas chalcedona kingstonensis; [iNaturalist] ; [BOA]
    • Occidryas chalcedona kingstonensis; [NACL], #4517e; [LepIndex]; [CoL]
  • E. c. trinitina Baughman & Murphy, 1998 California

    • Euphydryas chalcedona trinitina Baughman & Murphy, 1998; Sys. W. N. Am. Butts. (29): 400

    • Euphydryas (Euphydryas) chalcedona trinitina; Pelham, 2008, J. Res. Lepid. 40: 364; [P&N23]
    • Euphydryas chalcedona trinitina; [iNaturalist] ; [BOA]
• Euphydryas colon (Edwards, 1881) Washington, Oregon. See [maps] Colon Checkerspot

  • Melitaea colon Edwards, 1881; Papilio 1 (4): 45

  • Euphydryas (Euphydryas) colon colon; [P&N23]
  • Euphydryas (Euphydryas) colon; Pelham, 2008, J. Res. Lepid. 40: 365; [P&N23]
  • Euphydryas colon svihlae; [iNaturalist]
  • Euphydryas colon; [NAB], 605; [MPG] ; [iNaturalist] ; [BOA] ; [BOA] ; [BOA] ; [BOA] ; [BAMONA]
  • Lemonias colon; Dyar, 1903, Bull. U.S. nat. Mus. 52: 17
  • Occidryas colon; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [NACL], #4518; [LepIndex] ; [CoL]
  • ?Euphydryas chalcedona colon; [Opler]

  Larva on Symphoricarpos albus, Symphoricarpos vaccinoides, Penstemon antirrhinoides, P. subserratus, Plantago major [NAB]

  • E. c. colon

    • Melitaea perdiceas [= perdiccas] Edwards, 1881; Bull. Brookl. Ent. Soc. 3 (9): 80; TL: Mt Hood, Idaho
    • Melitaea perdiccas Edwards, 1881; Papilio 1 (11): 242 (emend. Melitaea periceas Edwars, 1881)
      • = ; Pelham, 2008, J. Res. Lepid. 40: 365
    • Euphydryas color tr.f. mcdunnoughi Gunder, 1928; Can. Ent. 60 (7): 163; TL: Corvallis, Oregon
      • = ; Pelham, 2008, J. Res. Lepid. 40: 366
    • Euphydryas chalcedona r. colon tr.f. fenderi Gunder, 1932; Pan-Pacific Ent. 8 (3): 125
      • = ; Pelham, 2008, J. Res. Lepid. 40: 366
    • Euphydryas colon ab. bakeri Fender, 1945; Bull. Brooklyn ent. Soc. 40 (2): 54 (preocc Stallings & Turner, 1945); TL: Elk Lake, Santiam Nat. Forest, Oregon
      • = ; Pelham, 2008, J. Res. Lepid. 40: 366

    • Euphydryas (Euphydryas) colon colon; Pelham, 2008, J. Res. Lepid. 40: 365
    • Euphydryas colon colon; [iNaturalist] ; [BOA]
    • Occidryas chalcedona perdiccas; [LepIndex]; [CoL]
    • Occidryas colon perdiccas; [NACL], #4518a
    • macdunnoughi[sic, mcdunnoughi]; dos Passos, 1964, Lepid. Soc. Mem. 1: 86
  • E. c. paradoxa McDunnough, 1927 British Colombia

    • Euphydryas perdiccas r. paradoxa McDunnough, 1927; Can. ent. 59 (7): 159; TL: Seton Lake, Lillooet, B. C:
    • Euphydryas perdiccas ab. ♂ nigrisupernipennis Gunder, 1926; Ent. News 37 (1): 4; TL: Chilcotin, British Columbia
      • = ; Pelham, 2008, J. Res. Lepid. 40: 367
    • Occidryas colon paradoxa; [NACL], #4518b

    • Euphydryas (Euphydryas) colon paradoxa; Pelham, 2008, J. Res. Lepid. 40: 366; [P&N23]
    • Euphydryas colon paradoxa; [iNaturalist] ; [BOA]
  • E. c. wallacensis Gunder, 1928 Idaho, ...

    • Euphydryas colon wallacensis Gunder, 1928; Pan-Pacific Ent. 5 (2): 50, pl. 2; TL: Wallace, Idaho
    • Euphydryas colon ab. huellemanni Comstock, 1926; Bull. South. Calif. Acad. Sci. 25 (1): 32; TL: Wallace, Idaho
      • = ; Pelham, 2008, J. Res. Lepid. 40: 366
    • Euphydryas chalcedona r. wallacensis tr.f. idahoensis Gunder, 1929; Pan-Pacific Ent. 6 (1): 3; TL: Wallace, Idaho
      • = ; Pelham, 2008, J. Res. Lepid. 40: 366

    • Euphydryas (Euphydryas) colon wallacensis; Pelham, 2008, J. Res. Lepid. 40: 366; [P&N23]
    • Euphydryas chalcedona wallacensis; [Wahlberg]
    • Euphydryas colon wallacensis; [iNaturalist] ; [BOA]
    • Occidryas chalcedona wallacensis; [LepIndex]; [CoL]
    • Occidryas colon wallacensis; [NACL], #4518c
  • E. c. sperryi Chermock & Chermock, 1945 California

    • Euphydryas colon sperryi Chermock & Chermock, 1945; Proc. Penn. Acad. Sci. 19: 39; TL: Bartle, California

    • Euphydryas (Euphydryas) colon sperryi; Pelham, 2008, J. Res. Lepid. 40: 365
    • Occidryas colon sperryi; [LepIndex]; [CoL]
  • E. c. nevadensis Bauer, 1975

    • Euphydryas colon nevadensis Bauer, 1975; Butterflies of North America: 181, pl. 42, f. 8, pl. 45, f. 11

    • Euphydryas (Euphydryas) colon nevadensis; Pelham, 2008, J. Res. Lepid. 40: 366; [P&N23]
    • Euphydryas chalcedona nevadensis; [Wahlberg]
    • Euphydryas colon nevadensis; [iNaturalist] ; [BOA]
    • Occidryas colon nevadensis; [NACL], #4518d
  • E. c. svilhae Pelham, 2008 Washington (Olympic Mtns)

    • Euphydryas (Euphydryas) colon svilhae Pelham, 2008; J. Res. Lepid. 40: 366; TL: Hurricane Hill, Olympic Mtns, Washington
    • Euphydryas chalcedona r. perdiccas tr.f. svilhae Gunder, 1932; Pan-Pacific Ent. 8 (3): 126; TL: Hurricane Hill, Olympic Mtns, Washington
• Euphydryas editha (Boisduval, 1852) -. See [maps] Edith's Checkerspot

  • Melitaea editha Boisduval, 1852; Ann. Soc. ent. Fr. (2) 10 (2): 304

  • Euphydryas (Euphydryas) editha editha; [P&N23]
  • Euphydryas (Euphydryas) editha; Pelham, 2008, J. Res. Lepid. 40: 357; [P&N23]
  • Euphydryas editha; [NAB], 607; [Opler]; [NL4A], #2063; [MPG] ; [iNaturalist] ; [BOA] ; [BOA] ; [BOA] ; [BOA] ; [BAMONA]
  • Lemonias editha; Dyar, 1903, Bull. U.S. nat. Mus. 52: 18
  • Occidryas editha; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [NACL], #4520; [LepIndex] ; [CoL]

  identification uncertain 1000x1290 (~179Kb) USA: Cabin Mountain, (1408m), (47°14'15N 121°16'27W), Kittitas Co., WA, 29.7.2006, Photo © Markku Savela

  identification uncertain 1000x1214 (~157Kb) USA: Cabin Mountain, (1408m), (47°14'15N 121°16'27W), Kittitas Co., WA, 29.7.2006, Photo © Markku Savela

  Larva on Schrophulariaceae , Castilleja , Orthocarpus , Pedicularis , Plantago [NAB]

  • E. e. editha

    • Euphydryas editha var. bayensis Sternitzky, 1937; Can. Ent. 69 (9): 204; TL: Hillsborough, San Mateo Co., Calif.
      • = ; Pelham, 2008, J. Res. Lepid. 40: 358
    • Euphydryas editha r. baroni tr.f. sternitzkyi Gunder, 1929; Pan-Pacific Ent. 6 (1): 6
      • = ; Pelham, 2008, J. Res. Lepid. 40: 358

    • Euphydryas (Euphydryas) editha editha; Pelham, 2008, J. Res. Lepid. 40: 358
    • Euphydryas editha bayensis; [iNaturalist]
    • Euphydryas editha editha; [iNaturalist] ; [BOA]
    • Occidryas editha bayensis; [NACL], #4520c
  • E. e. quino (Behr, 1863) California

    • Melitaea quino Behr, 1863; Proc. Calif. Acad. Sci. 3 (1): 90; TL: California
    • Melitaea augusta Edwards, 1890; Can. Ent. 22 (2): 21; TL: California
      • = ; [NL4A], #2063a; Pelham, 2008, J. Res. Lepid. 40: 358
    • Euphydryas editha ab. ♂ fieldi Gunder, 1924; Ent. News 35 (5): 155, pl. 2, f. E, D; TL: California
      • = ; [NL4A], #2063a; Pelham, 2008, J. Res. Lepid. 40: 358
    • Euphydryas editha r. wrighti Gunder, 1929; Pan-Pacific Ent. 6 (1): 5, pl. 9; TL: California
      • = ; [NL4A], #2063a; Pelham, 2008, J. Res. Lepid. 40: 358
    • Euphydryas editha wrighti tr.f. thornei Gunder, 1934; Can. Ent. 66 (6): 129; TL: California
      • = ; [NL4A], #2063a; Pelham, 2008, J. Res. Lepid. 40: 359
    • Euphydryas quino tr.f. rubyae Hower, 1935; Pan-Pacific Ent. 11 (2): 82; TL: California
      • = ; [NL4A], #2063a; Pelham, 2008, J. Res. Lepid. 40: 359
    • ?Occidryas kennei Gunder, 1932²

    • Euphydryas (Euphydryas) editha quino; Pelham, 2008, J. Res. Lepid. 40: 358; [P&N23]
    • Euphydryas chalcedona quino;
    • Euphydryas editha augusta; [BOW]: pl. 13, f. 24
    • Euphydryas editha quino; [Wahlberg]; [NL4A], #2063a; [iNaturalist] ; [BOA]
    • Lemonias augusta; Dyar, 1903, Bull. U.S. nat. Mus. 52: 18
    • Lemonias quino; Dyar, 1903, Bull. U.S. nat. Mus. 52: 18
    • Occidryas chalcedona quino; [NACL], #4517b; [LepIndex]; [CoL]
    • Occidryas editha augusta; [NACL], #4520n
    • Occidryas editha wrighti; [NACL], #4520a
  • E. e. nubigena (Behr, 1863) California

    • Melitaena nubigena Behr, 1863; Proc. Calif. Acad. Sci. 3 (1): 91
    • Euphydryas editha r. nubigena tr.f. tiogaensis Gunder, 1929; Pan-Pacific Ent. 6 (1): 6; TL: Tioga Pass, Yosemite Nat. Park, California
      • = ; Pelham, 2008, J. Res. Lepid. 40: 359
    • Euphydryas editha r. nubigena tr.f. boharti Gunder, 1929; Pan-Pacific Ent. 6 (1): 6; TL: Tuolumne Meadows, Yosemite Nat. Park, California
      • = ; Pelham, 2008, J. Res. Lepid. 40: 359

    • Euphydryas (Euphydryas) editha nubigena; Pelham, 2008, J. Res. Lepid. 40: 359; [P&N23]
    • Euphydryas editha nubigena; [iNaturalist] ; [BOA]
    • Euphydryas nubigena; [BOW]: pl. 13, f. 27
    • Lemonias nubigena; Dyar, 1903, Bull. U.S. nat. Mus. 52: 17
    • Occidryas editha nubigena; [NACL], #4520s; [LepIndex]; [CoL]
    • Occidryas tiogoensis[sic, tiogaensis]; McDunnough, 1938
    • nubigena; Mead, 1875
    • nubigeria[sic, nubigena]; Edwards, 1873
  • E. e. baroni (Edwards, 1879)

    • Melitaea baroni Edwards, 1879; Can. Ent. 11 (7): 129; TL: 'Mendocino'
    • Melitaea baroni H. Edwards, 1881; Papilio 1 (4): 52
    • Euphydryas edithar. baroni tr.f. dunni Gunder, 1929; Pan-Pacific Ent. 6 (1): 6; TL: Mendocino Co., California
      • = ; Pelham, 2008, J. Res. Lepid. 40: 360

    • Euphydryas (Euphydryas) editha baroni; Pelham, 2008, J. Res. Lepid. 40: 360; [P&N23]
    • Euphydryas editha baroni; [iNaturalist] ; [BOA]
    • Lemonias baroni; Dyar, 1903, Bull. U.S. nat. Mus. 52: 18
    • Occidryas editha baroni; [NACL], #4520j
  • E. e. rubicunda (H. Edwards, 1881) California

    • Melitaea quino var. rubicunda H. Edwards, 1881; Papilio 1 (4): 52
    • Euphydryas rubicunda tr.f. cottlei Gunder, 1928; Can. Ent. 60 (7): 164, pl. A, f. 7
      • = ; Pelham, 2008, J. Res. Lepid. 40: 360

    • Euphydryas (Euphydryas) editha rubicunda; Pelham, 2008, J. Res. Lepid. 40: 360; [P&N23]
    • Euphydryas editha rubicunda; [BOW]: pl. 13, f. 20; [iNaturalist] ; [BOA]
    • Lemonias rubicunda; Dyar, 1903, Bull. U.S. nat. Mus. 52: 18
    • Occidryas editha rubicunda; [NACL], #4520k
  • E. e. taylori (Edwards, 1888) British Columbia

    • Melitaea taylori Edwards, 1888; Can. Ent. 20 (5): 82
    • Euphydryas taylori ab. ♀ victoriae Gunder, 1926; 4, pl. 1, f. 6
      • = ; Pelham, 2008, J. Res. Lepid. 40: 358
    • Euphydryas taylori tr.f. barnesi Gunder, 1928; Can. Ent. 60 (7): 164, pl. A, f. 8, 8a
      • = ; Pelham, 2008, J. Res. Lepid. 40: 358

    • Euphydryas (Euphydryas) editha taylori; Pelham, 2008, J. Res. Lepid. 40: 358; [P&N23]
    • Euphydryas editha taylori; [iNaturalist] ; [BOA]
    • Euphydryas taylori; [BOW]: pl. 2, f. 6 (text only)
    • Lemonias taylori; Dyar, 1903, Bull. U.S. nat. Mus. 52: 18
    • Occidryas editha taylori; [NACL], #4520d; [LepIndex]; [CoL]

    1822x1234 (~544Kb) ♀ upperside Canada: British Columbia, Victoria, 6 May 1917 (topotype, W. Downes leg.), Photo 2007 © Norbert Kondla

    1772x1184 (~504Kb) ♀ underside Canada: British Columbia, Victoria, 6 May 1917 (topotype, W. Downes leg.), Photo 2007 © Norbert Kondla

  • E. e. beani (Skinner, 1897)

    • Melitaea beani Skinner, 1897; Can. Ent. 29 (6): 155
    • Euphydryas nubigena var. beani ab. ♂ blackmorei Gunder, 1926; Ent. News 37 (1): 2, pl. 1, f. 1
      • = ; Pelham, 2008, J. Res. Lepid. 40: 362

    • Euphydryas (Euphydryas) editha beani; Pelham, 2008, J. Res. Lepid. 40: 362; [P&N23]
    • Euphydryas editha beani; [iNaturalist] ; [BOA]
    • Lemonias anicia beani; Dyar, 1903, Bull. U.S. nat. Mus. 52: 17
    • Occidryas editha beani; [NACL], #4520c; [LepIndex]; [CoL]
  • E. e. augustina (Wright, 1905) S.California

    • Melitaea augusta var. augustina Wright, 1905; Butts. West Coast US: 154, pl. 19, f. 170

    • Euphydryas (Euphydryas) editha augustina; Pelham, 2008, J. Res. Lepid. 40: 358; [P&N23]
    • Euphydryas editha augustina; [iNaturalist] ; [BOA]
  • E. e. colonia (Wright, 1905) N.Oregon

    • Melitaea colonia Wright, 1905; Butts. West Coast US: 158, pl. 20, f. 180

    • Euphydryas (Euphydryas) editha colonia; Pelham, 2008, J. Res. Lepid. 40: 360; [P&N23]
    • Euphydryas colonia; [BOW]: pl. 2, f. 11 (text only)
    • Euphydryas editha colonia; [iNaturalist] ; [BOA]
    • Occidryas editha colonia; [NACL], #4520o; [LepIndex]; [CoL]
  • E. e. edithana Strand, 1915

    • Lemonias edithana Strand, 1915; Arch. Naturgesch. 80 A (11): 155

    • Euphydryas (Euphydryas) editha edithana; Pelham, 2008, J. Res. Lepid. 40: 361; [P&N23]
    • Euphydryas editha edithana; [iNaturalist] ; [BOA]
    • Occidryas editha edithana; [NACL], #4520i; [LepIndex]; [CoL]
  • E. e. aurilacus Gunder, 1928

    • Euphydryas aurilacus Gunder, 1928; Pan-Pacific Ent. 5 (2): 49, pl. 2, f. B; TL: nr Gold Lake Camp, Gold Lake, Siorra Co., California
    • Euphydryas rubicunda ab. ♂ foxi Gunder, 1924; Ent. News 35 (5): 155, pl. 2, f. D; TL: Gold Lake, Sierra Co., California
      • = ; Pelham, 2008, J. Res. Lepid. 40: 359
    • Euphydryas rubicunda ab. ♂ albiradiata Gunder, 1926; Ent. News 37 (1): 3, pl. 1, f. 4
      • = ; Pelham, 2008, J. Res. Lepid. 40: 359

    • Euphydryas (Euphydryas) editha aurilacus; Pelham, 2008, J. Res. Lepid. 40: 359; [P&N23]
    • Euphydryas editha aurilacus; [iNaturalist] ; [BOA]
    • Occidryas editha aurilacus; [NACL], #4520p; [LepIndex]; [CoL]
  • E. e. hutchinsi McDunnough, 1928 Montana

    • Euphydryas hutchinsi McDunnough, 1928; Can. ent. 60 (10): 248; TL: Milligan Canyon, Jefferson Co.

    • Euphydryas (Euphydryas) editha hutchinsi; Pelham, 2008, J. Res. Lepid. 40: 362; [P&N23]
    • Euphydryas editha hutchinsi; [iNaturalist] ; [BOA]
    • Occidryas editha hutchinsi; [NACL], #4520f; [LepIndex]; [CoL]
  • E. e. montanus McDunnough, 1928 Wyoming

    • Euphydryas hutchinsi var. montanus McDunnough, 1928; Can. ent. 60 (10): 249; TL: upper slopes Mt Washburn, Yellowstone Nat. park, 10000ft

    • Euphydryas (Euphydryas) editha montanus; Pelham, 2008, J. Res. Lepid. 40: 362; [P&N23]
    • Euphydryas editha montanus; [iNaturalist] ; [BOA]
  • E. e. monoensis Gunder, 1928

    • Euphydryas monoensis Gunder, 1928; Pan-Pacific Ent. 5 (2): 49, pl. 2, f. A
    • Euphydryas editha r. fridayi Gunder, 1931; Bull. South. Calif. Acad. Sci. 30 (2): 47
      • = ; Pelham, 2008, J. Res. Lepid. 40: 361
    • Euphydryas rubicunda ab. rubrosuffusa Comstock, 1926; Bull. South. Calif. Acad. Sci. 25 (1): 32; TL: Mammoth, Mono Co., California
      • = ; Pelham, 2008, J. Res. Lepid. 40: 361

    • Euphydryas (Euphydryas) editha monoensis; Pelham, 2008, J. Res. Lepid. 40: 361; [P&N23]
    • Euphydryas editha monoensis; [iNaturalist] ; [BOA]
    • Occidryas editha monoensis; [NACL], #4520l; [LepIndex]; [CoL]
  • E. e. lehmani Gunder, 1929

    • Euphydryas editha r. lehmani Gunder, 1929; Pan-Pacific Ent. 6 (1): 7, pl. 12; TL: Lehman Caves Resort, Mt Wheeler, White Pine Co., Nevada
    • Euphydryas editha lehmani tr.f. caverna Gunder, 1934; Can. Ent. 66 (6): 129; TL: Mt Wheele, White Pine Co., Nevada
      • = ; Pelham, 2008, J. Res. Lepid. 40: 361
    • Euphydryas editha koreti Murphy & Ehrlich, 1984; J. Res. Lepid. 22 (4): 257, 259
      • = ; Pelham, 2008, J. Res. Lepid. 40: 361

    • Euphydryas (Euphydryas) editha lehmani; Pelham, 2008, J. Res. Lepid. 40: 361; [P&N23]
    • Euphydryas editha lehmani; [iNaturalist] ; [BOA]
    • Occidryas editha lehmani; [NACL], #4520m; [LepIndex]; [CoL]
  • E. e. lawrencei Gunder, 1931 Oregon

    • Euphydryas editha r. lawrencei Gunder, 1931; Bull. South. Calif. Acad. Sci. 30 (2): 47
    • Euphydryas editha r. lawrencei tr.f. thielsenensis Gunder, 1931; Bull. South. Calif. Acad. Sci. 30 (2): 48
      • = ; Pelham, 2008, J. Res. Lepid. 40: 360
    • Euphydryas editha r. lawrencei tr.f. diamondensis Gunder, 1931; Bull. South. Calif. Acad. Sci. 30 (2): 48
      • = ; Pelham, 2008, J. Res. Lepid. 40: 360
    • Euphydryas editha remingtoni Burdick, 1959; Lepid. News 12 (5/6): 168
      • = ; Pelham, 2008, J. Res. Lepid. 40: 360

    • Euphydryas (Euphydryas) editha lawrencei; Pelham, 2008, J. Res. Lepid. 40: 359; [P&N23]
    • Euphydryas editha lawrencei; [BOA]
    • Occidryas editha lawrencei; [NACL], #4520r; [LepIndex]; [CoL]
    • Occidryas editha remingtoni; [NACL], #4520q
    • thielsonensis[sic, thielsenensis]; McDunnough, 1938
  • E. e. alebarki Ferris, 1971 Wyoming

    • Euphydryas editha alebarki Ferris, 1971; J. Res. Lepid. 9 (1): 17; TL: Carbon Co., Wyoming

    • Euphydryas (Euphydryas) editha alebarki; Pelham, 2008, J. Res. Lepid. 40: 362; [P&N23]
    • Euphydryas editha alebarki; [iNaturalist] ; [BOA]
    • Occidryas editha alebarki; [NACL], #4520g; [LepIndex]; [CoL]
  • E. e. gunnisonensis Brown, 1971

    • Euphydryas editha gunnisonensis Brown, 1971; J. Res. Lepid. 9 (1): 21; TL: Owl Creek Pass road 2 miles east of Colo. 550, nr Ridgeway, Ouray Co., Colorado, 7000ft

    • Euphydryas (Euphydryas) editha gunnisonensis; Pelham, 2008, J. Res. Lepid. 40: 361; [P&N23]
    • Euphydryas editha gunnisonensis; [iNaturalist] ; [BOA]
    • Occidryas editha gunnisonensis; [NACL], #4520h; [LepIndex]; [CoL]
  • E. e. insularis Emmel & Emmel, 1974

    • Euphydryas editha insularis Emmel & Emmel, 1974; J. Res. Lepid. 13 (2): 131; TL: California, Santa Barbara Co., Santa Rosa Is.

    • Euphydryas (Euphydryas) editha insularis; Pelham, 2008, J. Res. Lepid. 40: 358; [P&N23]
    • Euphydryas editha insularis; [iNaturalist] ; [BOA]
    • Occidryas editha insularis; [NACL], #4520b
  • E. e. luestherae Murphy & Ehrlich, 1980

    • Euphydryas editha luestherae Murphy & Ehrlich, 1980; J. Lep. Soc. 34 (3): 317; TL: Stanislaus Co., Del Puerto Canyon
    • Melitaea hoffmanni var. mirabilis Wright, 1905; Butts. West Coast US: 160, pl. 20, f. 184
      • = ; Pelham, 2008, J. Res. Lepid. 40: 360

    • Euphydryas (Euphydryas) editha luestherae; Pelham, 2008, J. Res. Lepid. 40: 360; [P&N23]
    • Euphydryas editha luestherae; [iNaturalist] ; [BOA]
  • E. e. ehrlichi Baughman & Murphy, 1998

    • Euphydryas editha ehrlichi Baughman & Murphy, 1998; Sys. W. N. Am. Butts. (29): 398

    • Euphydryas (Euphydryas) editha ehrlichi; Pelham, 2008, J. Res. Lepid. 40: 359; [P&N23]
    • Euphydryas editha ehrlichi; [iNaturalist] ; [BOA]
  • E. e. karinae Baughman & Murphy, 1998

    • Euphydryas editha karinae Baughman & Murphy, 1998; Sys. W. N. Am. Butts. (29): 399

    • Euphydryas (Euphydryas) editha karinae; Pelham, 2008, J. Res. Lepid. 40: 360; [P&N23]
    • Euphydryas editha karinae; [iNaturalist] ; [BOA]
  • E. e. bingi Baughman & Murphy, 1998

    • Euphydryas editha bingi Baughman & Murphy, 1998; Sys. W. N. Am. Butts. (29): 399; TL: California, Modoc Co., Warner Mts., Emerson Peak, 8200ft

    • Euphydryas (Euphydryas) editha bingi; Pelham, 2008, J. Res. Lepid. 40: 361; [P&N23]
    • Euphydryas editha bingi; [iNaturalist] ; [BOA]
  • E. e. tahoensis Austin & Murphy, 1998

    • Euphydryas editha tahoensis Austin & Murphy, 1998; in Emmel, Systematics W. North Am. Butts.: 408; TL: Nevada, Washoe Co.; Carson Range, Tahoe Meadows, 2590m

    • Euphydryas (Euphydryas) editha tahoensis; [P&N23]
    • Euphydryas editha tahoensis; [iNaturalist] ; [BOA]
  • E. e. owyheensis Austin & Murphy, 1998

    • Euphydryas editha owyheensis Austin & Murphy, 1998; in Emmel, Systematics W. North Am. Butts.: 410; TL: Nevada, Elko Co.

    • Euphydryas (Euphydryas) editha owyheensis; Pelham, 2008, J. Res. Lepid. 40: 361; [P&N23]
    • Euphydryas editha owyheensis; [iNaturalist] ; [BOA]
  • E. e. mattooni Austin & Murphy, 1998

    • Euphydryas editha mattooni Austin & Murphy, 1998; in Emmel, Systematics W. North Am. Butts.: 411; TL: California, Modoc Co.

    • Euphydryas (Euphydryas) editha mattooni; Pelham, 2008, J. Res. Lepid. 40: 361; [P&N23]
    • Euphydryas editha mattooni; [iNaturalist] ; [BOA]

• Euphydryas maturna (Linnaeus, 1758) NW.China, Mongolia. See [maps] Broknätfjäril Eschenscheckenfalter Przeplatka maturna Scarce Fritillary Kleiner Maivogel Kirjoverkkoperhonen

  • Papilio maturna Linnaeus, 1758; Syst. Nat. (Edn 10) 1: 480; TL: Sweden
  • Papilio cynthia Esper, 1778; Die Schmett. Th. I, Bd. 1 (9): 347, (7): pl. 37, f. 2
  • Papilio agrotera Bergsträsser, 1780; Nomen. Ins. 4: 13
  • Melitaea maturna var. urbani Hirschke, 1901; //www.zobodat.at/pdf/JBWEV_11_0085.pdf] Jber. Wien. ent. Ver. 11: 85
  • Melitaea maturna f. schlumbergeri Schultz, 1907; Ent. Zs. 20: 279
  • Melitaea maturna ab. fulgida Schultz, 1908; Ent. Zs. 22 (23): 90; TL: Wien
  • Melitaea maturna ab. kautzi Schawerda, 1913; Verh. zool.-bot. Ges. Wien 63: 146
  • Melitaea maturna ab. diabolus Schawerda, 1914; Verh. zool.-bot. Ges. Wien 64 (9/10): 352
  • pauper (Caruel, 1944) (Melitaea); Misc. ent. 41: 9
  • unifasciata (Caruel, 1944) (Melitaea); Misc. ent. 41: 9
  • arcuata (Caruel, 1944) (Melitaea); Misc. ent. 41: 10
  • fulvuniformis Verity, 1950; Le Farfalle diurn. d'Italia 4: 86

  • Euphydryas (Hypodryas) maturna; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 37
  • Euphydryas maturna; [SPRK]; [H&R], 125; [BOW]: pl. 2, f. 11; [EBW]; [BARU], #152; [BRU2]: 57, pl. 37, f. 16-18; [Wahlberg]; [FE]; [Otakar Kudrna]; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE] ; [SVF] ; [Mario Maier]
  • Hypodryas maturna; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [LepIndex] ; [CoL]

  Misidentified/misapplied:

  • ?Papilio maturna; Denis & Schiffermüller, 1775, Ank. syst. Schmett. Wienergegend: 179

  800x612 (~60Kb) ♂ upperside FINLAND: Kb: Kiihtelysvaara, 692:68, 1.7.1998 Photo © Markku Savela	1000x765 (~67Kb) underside FINLAND: Sa: Ristiina, 6817:528, 2.7.1998, Photo © Markku Savela	594x400 (~49Kb) larva Central Europe 1990-2000, Photo © Michael Borsch
  595x400 (~62Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch	527x511 (~113Kb) Russia, Moscow area, 4.7.2015 (36°25'E, 56°00'N), Photo © D. Smirnov	635x618 (~105Kb) Russia, Moscow area, 11.6.2011 (38°52'E, 55°33'N), Photo © D. Smirnov

  Larva on Betula spp., Salix spp., Fraxinus excelsior, Veronica longifolia, Melampyrum sylvaticum, Viburnum opulus, Lonicera xylosteum, Lonicera coerulea, L. caprifolium [SPRK], Scabiosa , Veronica , Salvia , Plantago , Viola , Digitalis [BRU2]: 58

  • E. m. maturna NEU, CEU

    • Euphydryas maturna maturna; [BRU2], 57; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 37
  • E. m. staudingeri Wnukowsky, 1929 Altai, Siberia, Transbaikalia, Dzhungarski Alatau, Mongolia?, NW.China?

    • maturna staudingeri Wnukowsky, 1929; Zool. Anz. 83: 121 (repl. Melitaea maturna var. uralensis Staudinger, 1871)
    • Melitaea maturna var. uralensis Staudinger, 1871; Cat. Lepid. eur. Faunegeb. (Edn. 2): 17 (preocc.)
    • Melitaea maturna idunides Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 3; TL: Bosnia, Maklenpass

    • Euphydryas maturna staudingeri; [BARU](note) , ; [BRU2], 57; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 37; [iNaturalist]
    • Hypodryas maturna staudingeri; [LepIndex]; [CoL]
    • Melitaea maturna var. uralensis; Staudinger, 1881, Stettin ent. Ztg 42 (7-9): 287; Staudinger, 1892, Dt. Ent. Z. Iris 5 (2): 321

    392x393 (~14Kb) ♀ underside A hillock swamped meadow at a riparian spruce wood at the vllage Topuchaya, Shebalino District, Altai Republic, the Altai Mts., S Siberia, Russia. 8th July 1998 (on Pentaphylloides fruticosa), Photo © Oleg Kosterin	480x408 (~41Kb) ♂ upperside A forb meadowy edge of a birch forest, the lower part of the Kora River valley, the Mynchukur mountain range, the northern chain of the Dzhungarian Alatau Mts., Taldy-Kurgan Province, SE Kazakhstan. 13th June 1994, Photo © Oleg Kosterin	1072x790 (~197Kb) ♂ upperside Kuzuyak Pass between the lower valleys of Akkem and Kucherla, the Katunskii Range northern principal slope, Central [Russian} Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 26th June 2007 (on Carum carvi), Photo © Oleg Kosterin
    1146x1065 (~260Kb) ♂ upperside Kuzuyak Pass between the lower valleys of Akkem and Kucherla, the Katunskii Range northern principal slope, Central [Russian} Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 26th June 2007 (on Carum carvi), Photo © Oleg Kosterin	1078x1042 (~277Kb) ♂ upperside Russia, West Siberia, NW Altai Mts., Ust'-Kan District, Bashchelakskii Range, meadow steppe 1 km SSW of Keley village. 4th July 2005, Photo © Oleg Kosterin	1235x1002 (~385Kb) ♂ upperside Russia, West Siberia, NW Altai Mts., Ust'-Kan District, Bashchelakskii Range, meadow steppe 1 km SSW of Keley village. 4th July 2005, Photo © Oleg Kosterin

  • E. m. aestia (Bois-Reymond, 1931) Germany

    • maturna aestia (Bois-Reymond, 1931) (Melitaea); Zs. wiss. InsekBiol. 26: 34

    • Hypodryas maturna aestia; [LepIndex]; [CoL]
  • E. m. urbanoides (Warnecke, 1942) Germany

    • maturna urbanoides (Warnecke, 1942) (Melitaea); Dt. ent. Z. Iris 56: 103

    • Hypodryas maturna urbanoides; [LepIndex]; [CoL]
  • E. m. coreanica Murayama, 1955 Korea

    • Euphidryas[sic] maturna coreanica Murayama, 1955; Tyô to Ga 6 (1): 1, f. 4; TL: Kosho, Heian-kokude, N-Korea

    • Hypodryas maturna coreanica; [LepIndex]; [CoL]
  • E. m. adamczewskii Krzywicki, 1967 Poland

    • Euphydryas maturna adamczewskii Krzywicki, 1967; Ann. zool. Warsz. 25 (1): 113

    • Hypodryas maturna adamczewskii; [LepIndex]; [CoL]
  • E. m. macedonica Dufay, 1977 Yugoslavia

    • maturna macedonica Dufay, 1977; Entomops 41: 28

    • Hypodryas maturna macedonica; [LepIndex]; [CoL]
  • E. m. pseudomaturna Lang, 2010 Heilongjiang

    • Euphydryas maturna pseudomaturna Lang, 2010; Atalanta 41: 223; TL: Heilongjiang, Jiamusi, Liangzihe
• Euphydryas italica Back, Haussmann, Salk & Weiss, 2015 N.Italy (Cuneo). See [maps]

  • Euphydryas italica Back, Haussmann, Salk & Weiss, 2015; Atalanta 46: 112; TL: Italy, "Cottische Alpen, Umebung Cuneo", 1000m

  • Euphydryas italica; [iNaturalist] ; [LEPIFORUM]
• Euphydryas intermedia (Ménétriés, 1859) -. See [maps] Asian Fritillary

  • Melitaea intermedia Ménétriés, 1859; in Schrenck, Reise Forschungen Amur-Lande 2 (1): 22; TL: Amurland

  • Euphydryas (Hypodryas) intermedia; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 37
  • Euphydryas intermedia; [H&R], 125; [BOW]: pl. 2, f. 11 (text only); [BRU2], 58; [MRS], 585; [Wahlberg]; [FE]; [Otakar Kudrna]; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE]
  • Hypodryas intermedia; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [LepIndex] ; [CoL]

  547x400 (~46Kb) ♂ upperside Dwarf pine/dwarf alder forest-tundra at about 1000 m above sea level, the mountains of the Koni Peninsula, Ola District, S Magadan Province, Russia. 23rd July 1989 (on Ledum palustre), Photo © Oleg Kosterin	878x932 (~273Kb) ♂ upperside A large opening with a ruderal vegetation, called Konoplyanaya Polyana, in the lower reaches of the Oroktoi brook, a right tributary of the Akkem River, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 27th June 2007 (on Ferula soongarica), Photo © Oleg Kosterin	1225x1110 (~356Kb) ♀ upperside An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 3rd July 2007 (on Patrinia rupestris), Photo © Oleg Kosterin
  915x1176 (~353Kb) ♀ upperside An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 4th July 2007, Photo © Oleg Kosterin	831x961 (~190Kb) ♂ underside An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 5th July 2007 (on Bistorta major), Photo © Oleg Kosterin	1201x872 (~417Kb) ♀ upperside The lower Oroktoi brook valley, a right tributary of the Akkem River, the Katunskii Range northern principal slope, Central [Russian} Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 7th July 2007 (on fresh horse dung), Photo © Oleg Kosterin

  Larva on Veronica , Lonicera [RFEL]

  • E. i. intermedia S.Siberia, Transbaikalia, Far East, Amur, Ussuri

    • Euphydryas intermedia intermedia; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 37; [iNaturalist]
  • E. i. wolfensbergeri (Frey, 1880) Alps (Maritime Alps)

    • Melitaea maturna var. wolfensbergeri Frey, 1880; Lep. Schweiz.: 27
      • = Euphydryas intermedia; [FE]

    • Euphydryas ichnea wolfensbergeri; [BRU2], 58
    • Euphydryas intermedia wolfensbergeri; [H&R], 125; [iNaturalist]
    • Hypodryas intermedia wolfensbergeri; [LepIndex]; [CoL]
  • E. i. mongolica (Staudinger, 1892) Sayan

    • Melitaea maturna ab. mongolica Staudinger, 1892; Dt. Ent. Z. Iris 5 (2): 321; TL: ["Kentei-gebirge"]

    • Euphydryas ichnea mongolica; [BRU2], 58
    • Euphydryas intermedia mongolica; [MRS], 585
  • E. i. altaiana (Wnukowsky, 1929) Altai

    • intermedia altaiana (Wnukowsky, 1929) (Melitaea); Zool. Anz. 83: 122 (repl. altaica Bang-Haas, 1909)
    • Melitaea maturna altaica Bang-Haas, 1909; Gross-Schmett. Erde 1: 212, pl. 65 b (preocc.)

    • Euphydrays ichnea altaica; [BARU](note) ,
    • Euphydryas ichnea altaiana; [BRU2], 58
    • Euphydryas intermedia altaica;

    906x600 (~84Kb) ♀ upperside Dwarf birch (Betula rotundifolia) thickets on the left bank of the Dzhazator River in its upper flow, between the falls of its right tributaries Chikty and Akbul, 2000 m above sea level, Kosh-Agach District, Altai Republic, West Siberia, Russia. 19th July 1998, Photo © Oleg Kosterin

  • E. i. konumensis (Matsumura, 1927) Sakhalin

    • Melitaea maturna konumensis Matsumura, 1927; Ins. Matsumurana 2 (2): 117, pl. 3, f. 5; TL: Saghalien

    • Euphydryas ichnea konumensis; [BRU2], 58
    • Euphydryas intermedia konumensis; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 37
  • E. i. janica (Lastukhin, 2008) N.Yakutia

    • intermedia janica (Lastukhin, 2008) (Hypodryas)

    • Euphydryas intermedia janica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 37
• Euphydryas cynthia ([Schiffermüller], 1775) Alps, Bulgaria. See [maps] Cynthia's Fritillary Veilchen-Scheckenfalter

  • Papilio cynthia [Schiffermüller], 1775; Ank. syst. Schmett. Wienergegend: 179
  • Papilio[?] trivia Esper, 1781; Die Schmett. Th. I, Bd. 2 (2): pl. 60, f. 1
  • Papilio mysia Hübner, [1799-1800]; Samml. eur. Schmett. [1]: pl. 1, f. 3
  • Papilio maturna Hübner, [1805]; Samml. eur. Schmett. [1]: pl. 117, f. 598-599
  • Melitaea cynthia ab. leucophryne Ochsenheimer, 1816; Schmett. Eur. 4: 103
  • Melitaea cynthia ab. impunctata Hoffmann, 1911; Int. ent. Zs. 5 (32): 227
  • Melitaea cynthia ab. brunneomarginata Kiefer, 1912; Ent. Rdsch. 29: 70
  • aurinoides (Kiefer, 1918) (Melitaea); Z. Öst. EntVer. 3: 108
  • Melitaea cynthia ab. obscurior Hofer, 1920; Soc. ent. 35: 46
  • Melitaea cynthia ab. tricolor Schawerda, 1926; Verh. zool.-bot. Ges. Wien 74-75 (s.b): 67
  • latevittata (Pionneau, 1936) (Melitaea); Echange 52: 24

  • Euphydryas cynthia; [H&R], 126; [BOW]: pl. 2, f. 8; [EBW]; [Wahlberg]; [FE]; [Otakar Kudrna]; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE]
  • Hypodryas cynthia; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [LepIndex] ; [CoL]

  Larva on Plantago , P. alpina, Alchemilla [H&R]

  • E. c. cynthia

    • Euphydryas cynthia cynthia; [H&R], 126
  • E. c. pallida (Spuler, 1901)

    • Melitaea cynthia pallida Spuler, 1901; Schmett. Eur. 1: 21

    • Melitaea cynthia pallida; Fruhstorfer, 1910, Soc. ent. 25 (13): 52
  • E. c. variegata (Turati & Verity, 1911)

    • Melitaea cynthia pallida f. variegata Turati & Verity, 1911; Boll. Soc. Ent. Ital. 42 : 205

    • Hypodryas cynthia variegata; [LepIndex]; [CoL]
  • E. c. mundata (Turati & Verity, 1911)

    • Melitaea cynthia pallida f. mundata Turati & Verity, 1911; Boll. Soc. Ent. Ital. 42 : 206

    • Hypodryas cynthia mundata; [LepIndex]; [CoL]
  • E. c. reducta (Vorbrodt, 1911) Switzerland

    • Melitaea cynthia f. reducta Vorbrodt, 1911; Die Schmett. Schweiz 1: 47

    • Hypodryas cynthia reducta; [LepIndex]; [CoL]
  • E. c. leonhardi (Fruhstorfer, 1917) Bulgaria

    • Melitaea cynthia leonhardi Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 5, pl. 1, f. 2-3; TL: Bulgaria, Rhodope
    • drenowskii (Röber, 1926) (Melitaea); Ent. Mitt. 15 (3/4): 227

    • Euphydryas aurinia drenowskii; [iNaturalist]
    • Hypodryas cynthia leonhardi; [LepIndex]; [CoL]
  • E. c. alpicola (Galvagni, 1918) Austria

    • Melitaea cynthia v. alpicola Galvagni, 1918; Jb. Wien. Ent. Vre. 28: 134
      • = Euphydryas cynthia; [FE]

    • Euphydryas cynthia alpicola; [H&R], 126
    • Hypodryas cynthia alpicola; [LepIndex]; [CoL]
  • E. c. umbrata (Galvagni, 1918)

    • Melitaea cynthia ab. umbrata Galvagni, 1918; Jb. Wien. Ent. Vre. 28: 136

    • Hypodryas cynthia umbrata; [LepIndex]; [CoL]
  • E. c. fasciata (Wehrli, 1920)

    • cynthia fasciata (Wehrli, 1920) (Melitaea); Mitt. ent. Ver. Basel.: 2

    • Hypodryas cynthia fasciata; [LepIndex]; [CoL]
  • E. c. privata (Stauder, 1922)

    • Melitaea cynthia aurinoides f. privata Stauder, 1922; Ent. Anz. 2 (3): 30

    • Hypodryas cynthia privata; [LepIndex]; [CoL]
  • E. c. sterlineata (Turati, 1923) Italy

    • cynthia sterlineata (Turati, 1923) (Melitaea); Mem. Soc. Ent. 2: 6

    • Hypodryas cynthia sterlineata; [LepIndex]; [CoL]
  • E. c. spuleri (Röber, 1926) Bulgaria

    • cynthia spuleri (Röber, 1926) (Melitaea); Ent. Mitt. 15 (3/4): 226

    • Hypodryas cynthia spuleri; [LepIndex]; [CoL]
  • E. c. extensa (Dujardin, 1948)

    • cynthia extensa (Dujardin, 1948) (Melitaea); Bull. Soc. ent. Mulhouse 4 (1948): 16

    • Hypodryas cynthia extensa; [LepIndex]; [CoL]
  • E. c. minor Verity, 1950 Italy

    • cynthia minor Verity, 1950; Le Farfalle diurn. d'Italia 4: 81

    • Hypodryas cynthia minor; [LepIndex]; [CoL]
  • E. c. demissa Verity, 1950 Italy

    • cynthia demissa Verity, 1950; Le Farfalle diurn. d'Italia 4: 83

    • Hypodryas cynthia demissa; [LepIndex]; [CoL]
• Euphydryas iduna (Dalman, 1816) . See [maps] Lappnätfjäril Lapinverkkoperhonen Lapland Fritillary

  • Melitaea iduna Dalman, 1816; K. VetenskAcad. Handl. 1816 (1): 75; TL: Sweden
  • Melitaea ichnea Boisduval, [1833]; Icon. hist. Lépid. Europe 1 (11-12): 112, pl. 23, f. 5-6; TL: "le nord de la Laponiae et de la Sibirie"
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 47
  • Melitaea iduna ab. sulitelmica Schultz, 1904; Abh. Nat. Ges. Görlitz 24: 137; TL: Sulitelma
  • Melitaea iduna ab. nigrofasciata Lingonblad, 1944; Notulae Ent. 24 (1-2) (1-2): 49; TL: Finland, Kilpisjärvi
  • Melitaea iduna ab. saanae Lingonblad, 1944; Notulae Ent. 24 (1-2) (1-2): 49; TL: Finland, Kilpisjärvi

  • Euphydrya iduna; [SPRK]; [H&R], 127
  • Euphydryas (Hypodryas) iduna; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 37
  • Euphydryas ichnea ichnea; [BRU2], 58
  • Euphydryas ichnea; [BARU], #153; [BRU2]: 58, pl. 37, f. 19-21; Yakovlev, 2012, Nota lepid. 35 (1): 87
  • Euphydryas iduna; [BOW]: pl. 2, f. 10; [EBW]; [BARU], #154; [MRS], 585; [BRU2]: 58, pl. 37, f. 22-30; [Wahlberg]; [FE]; [Otakar Kudrna]; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 47; Yakovlev, 2012, Nota lepid. 35 (1): 87; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE] ; [SZMN] ; [SVF] ; [Mario Maier]
  • Hypodryas iduna; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [LepIndex] ; [CoL]

  400x520 (~50Kb) pupa Finland, Kilpisjärvi 767:25, 06.07.1982, Photo © Pertti Pakkanen

  Larva on (as ichnea) Lonicera maacki [BRU2]: 58, Bartsia alpina?, Vaccinium uliginosum?, V. myrtillus? [SPRK], Plantago , Veronica , Vaccinium [BRU2]: 59

  • E. i. iduna Arctic Europe, Arctic Asia

    • Euphydryas iduna iduna; [BRU2], 58; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 37
  • E. i. sajana Higgins, 1950 Altai, Sayan

    • iduna sajana Higgins, 1950; Trans. R. ent. Soc. Lond. 101 (12): 443
    • sajana (Bang-Haas, 1930) (Melitaea); Gross-Schmett. Erde 1 (Suppl.): 204
    • sajana Gaede, 1932; (nom. nud.)

    • Euphydryas iduna sajana; [BRU2], 58; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 37; Yakovlev, 2012, Nota lepid. 35 (1): 87
  • E. i. inexpectata (Sheljuzhko, 1934) Caucasus Major

    • Melitaea iduna inexpectata Sheljuzhko, 1934; Zs. Öst. EntVer. 19: 30; TL: Chatipara Mtns, 2700m

    • Euphydryas iduna inexpectata; [BRU2], 58; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 47; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 37
    • Hypodryas iduna inexpectata; [LepIndex]; [CoL]
  • E. i. alferakyi Korshunov, 1996 Far East

    • Euphydryas iduna alferakyi; [Oleg Kosterin]; [BRU2], 58; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 37; [SZMN]

    500x447 (~92Kb) ♀ upperside A tremendows large-stoned scree at about 1000 m above sea level, the mountains of the Koni Peninsula, Ola District, S Magadan Province, Russia. 23rd July, 1989, Photo © Oleg Kosterin

    533x400 (~58Kb) underside A subalpine meadow at the dwarf pine (Pinus pumila (Pall.) Regel) thickets on the southen slope of a pass between Khindzha and Burgauli valleys, in a valley of a left tributary of the latter, about 500 m above sea level, the mountains of the Koni Peninsula, Ola District, S Magadan Province, Russia. 12th July, 1989, Photo © Oleg Kosterin

  • E. i. semenovi Korshunov & Ivonin, 1996 Kuznetsky Alatau Mts.

    • Euphydryas iduna semenovi; [BRU2], 58; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 37; [SZMN]
  • E. i. gorodinskii Churkin & Kolesnichenko, 2003

    • Euphydryas iduna gorodinskii Churkin & Kolesnichenko, 2003; Helios 4: (92-119); TL: Russia, Kamchatka, Bystrinskyi distr., Esso v., Uksichan R.

    • Euphydryas iduna gorodinskii; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38
  • E. i. emerita Churkin & Kolesnichenko, 2003 Altai, Sayan

    • Euphydryas iduna emerita Churkin & Kolesnichenko, 2003; Helios 4: (92-119); TL: South Mongolia, Gobi-Altai Aimak, 30km S fr Biger somon

    • Euphydryas iduna emerita; Yakovlev, 2012, Nota lepid. 35 (1): 87; [iNaturalist]

    942x600 (~88Kb) upperside An alpine meadow in the valley of the left headwater of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 10th July 1998, Photo © Oleg Kosterin	457x500 (~42Kb) underside Highland dwarf birch (Betula rotundifolia) tundras on a ledge of the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2300 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 15th July 1998, Photo © Oleg Kosterin	864x1067 (~236Kb) ♂ upperside An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 3rd July 2007 (on Bupleurum), Photo © Oleg Kosterin
    1080x863 (~433Kb) ♀ upperside An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 3rd July 2007, Photo © Oleg Kosterin	1115x918 (~294Kb) ♀ upperside An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 3rd July 2007, Photo © Oleg Kosterin

  • E. i. putorana (Lastukhin, 2008)

    • iduna putorana (Lastukhin, 2008) (Hypodryas)
    • tamara Lastuksin, 2008
      • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 37

    • Euphydryas iduna putorana; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 37
  • E. i. jacobsoni Yakovlev, 2011 E.Kazahstan

    • Euphydryas iduna jacobsoni Yakovlev, 2011; Atalanta 42 (1-4): 227; TL: E.Kazakhstan, Saur Mts, upper stream of B. Dzhemenej river, 2150m
• Euphydryas gillettii (Barnes, 1897) Wyoming. See [maps] Gillette's Checkerspot

  • Melitaea gillettii Barnes, 1897; Can. Ent. 29 (2): 40; TL: Yellowstone Park, Wyoming
  • Melitaea glacialis Skinner, 1921; Ent. News 32 (3): 89; TL: Montana
    • = ; Pelham, 2008, J. Res. Lepid. 40: 357
  • Euphydryas gilletti tr.f. herri Gunder, 1929; Pan-Pacific Ent. 6 (1): 8; TL: Martina, near Stark, Missoula Co., Montana
    • = ; Pelham, 2008, J. Res. Lepid. 40: 357

  • Euphydryas (Hypodryas) gillettii; Pelham, 2008, J. Res. Lepid. 40: 357; [P&N23]
  • Euphydryas gillettii; [NAB], 608; [Wahlberg]; [Opler]; [MPG] ; [iNaturalist] ; [BOA] ; [BAMONA]
  • Hypodryas gillettii; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [NACL], #4521; [LepIndex] ; [CoL]
  • Lemonias gillettei[sic, gillettii]; Dyar, 1903, Bull. U.S. nat. Mus. 52: 17
  • gracialis[sic, glacialis]; McDunnough, 1938

  Larva on Lonicera involucrata, Symphoricarpos albus [NAB]

• Euphydryas aurinia (Rottemburg, 1775) Yakutia, NW.China, Mongolia. See [maps] Punakeltaverkkoperhonen Przeplatka aurinia Ärenprisnätfjäril Marsh Fritillary Guldnätfjäril Goldener Scheckenfalter Ondas rojas

  • Papilio aurinia Rottemburg, 1775; Der Naturforscher 6: 5; TL: Paris
  • Papilio artemis Denis & Schiffermüller, 1775; Ank. syst. Schmett. Wienergegend: 322 (preocc.)
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 46
  • Papilio maturna Esper, 1777; Die Schmett. Th. I, Bd. 1 (8): 209, (3): pl. 16, f. 2 (preocc.)
  • ?Papilio maturna Esper, 1778; Die Schmett. Th. I, Bd. 1 (8): 209, (7): pl. 37
  • Papilio lye Bergsträsser, 1780; Nomen. Ins. 5[?]: pl. 82, f. 4-5
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 46
  • Papilio koloswarensis Piller & Mitterpacher, 1783; Iter Poseganam: 41
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 46
  • Papilio matutina Thunberg, 1791; Diss. Ent. sistens Insecta Suecica (2): 45
  • f. minima Gianelli, 1890; Ann. Acc. Agric. Torino 33: 19
  • Melitaea aurinia ab. dubia Krulikovsky, 1890; Bull. Soc. imp. Nat. Moscou (n.s.) 4 (2): 234, pl. 8, f. f
  • Melitaea aurinia ab. obscurata Krulikovsky, 1890; Bull. Soc. imp. Nat. Moscou (n.s.) 4 (2): 235, pl. 8, f. c
  • Melitaea aurinia ab. provincialis Tutt, 1896; Brit. Butts: 316
  • Melitaea aurinia ab. brunnea Tutt, 1896; Brit. Butts: 316
  • Melitaea aurinia ab. ochrea Tutt, 1896; Brit. Butts: 318
  • Melitaea aurinia ab. virgata Tutt, 1896; Brit. Butts: 318
  • Melitaea aurinia ab. nigrolimbata Schultz, 1906; Ent. Zs. 19 (35): 205
  • Melitaea aurinia ab. atricolor Schultz, 1906; Ent. Zs. 19 (35): 205
  • Melitaea aurinia ab. impunctata Schultz, 1906; Ent. Zs. 19 (35): 205
  • Melitaea aurinia ab. sesostris Schultz, 1906; Ent. Zs. 19 (35): 206
  • Melitaea aurinia ab. sebaldus Schultz, 1906; Ent. Zs. 19 (35): 206
  • nana (Rehfons, 1910) (Melitaea); Cat. Rhop. Genève: 12
  • Militaea[sic] aurinia ab. fulvacea Lambillion, 1911; Rev. mens. Soc. ent. Namur. 11: 78
  • turbida (Gélin & Lucas, 1912) (Melitaea); Mem. Soc. deux. Sevres 1912: 34
  • unipuncta (Kiefer, 1916) (Melitaea); Ent. Jb. 28: 134
  • hoffmanni (Kiefer, 1916) (Melitaea); Ent. Jb. 28: 134
  • Melitaea aurinia f. merope-minor Vorbrodt & Müller-Rutz, 1917; Mitt. schweiz. ent. Ges. 12 (9-10): 435 (Krüger i.l.)
  • Melitaea iberica ab. nigrolunulata Fernández, 1918; Bol. Soc. Ent. España 1 (8): 158; TL: Sierra de Alfacar
  • semifuscata (Cabeau, 1919) (Melitaea); Rev. mens. Soc. ent. Namur. 1919: 61
  • epimolpadia (Reverdin, 1919) (Melitaea); Bull. Soc. Lep. Genève 4: 108
  • Melitaea aurinia comacina ab. sterlineata Turati, 1921; Naturalista sicil. 23 (7-12): 221
  • Melitaea aurinia comacina ab. denigrata Turati, 1921; Naturalista sicil. 23 (7-12): 221
  • didyma (Heinrich, 1923) (Melitaea); Dt. ent. Zs. 1923 (Beiheft): 48
  • rectiangula (Cabeau, 1924) (Melitaea); Rev. mens. Soc. ent. Namur. 24: 17
  • Melitaea aurinia v. iberica ab. demaculata Bubacek, 1923; Verh. zool.-bot. Ges. Wien 73 (s.b): 24
  • catherini (Le Charles, 1924) (Melitaea); Amat. Papillons 2: 88
  • perianthes (Cabeau, 1926) (Melitaea); Lambillionea 26:74
  • interligata (Derenne, 1926) (Melitaea); Lambillionea 26: 91
  • gracilens (Cabeau, 1927) (Melitaea); Lambillionea 27: 10
  • geminifasciata (Cabeau, 1927) (Melitaea); Lambillionea 27: 11
  • transversa (Cabeau, 1928) (Melitaea); Lambillioena 28: 11
  • deficiens (Cabeau, 1928) (Melitaea); Lambillioena 28: 11
  • leucophana (Cabeau, 1928) (Melitaea); Lambillionea: 78
  • f. translucens (Verity, 1928) (Melitaea); Ent. Rec. 40 (3): 43
  • frigescens (Verity, 1928) (Melitaea); Ent. Rec. 40 (3): 45
  • moritura (Verity, 1928) (Melitaea); Ent. Rec. 40 (6): 89
  • debilis-provincialis [= debilisprovincialis] (Verity, 1928) (Melitaea); Ent. Rec. 40 (6): 90
  • semigriseis (Cabeau, 1931) (Melitaea); Lambillionea 31: 3
  • tetramelana (Cabeau, 1931) (Melitaea); Lambillionea 31: 174
  • semigracilens (Cabeau, 1931) (Melitaea); Lambillionea 31: 199
  • melanoleuca (Cabeau, 1932) (Melitaea); Lambillionea 32: 76
  • flavofasciata (Hackray, 1934) (Melitaea); Lambillionea 34: 126
  • insterburgia (Braun, 1937) (Melitaea); Ent. Rundsch. 54: 556
  • albofasciata Frohawk, 1938; Var. Brit. Butterfl. : pl. 17, f. 1
  • suffusa Frohawk, 1938; Var. Brit. Butterfl. : pl. 17, f. 3
  • infra-rectiangula (Caruel, 1939) (Melitaea); Lambillionea 39: 46
  • ocellata (Caruel, 1944) (Melitaea); Misc. ent. 41: 9
  • arcuata (Caruel, 1944) (Melitaea); Misc. ent. 41: 10
  • commacula (Caruel, 1944) (Melitaea); Misc. ent. 41: 10
  • ligata (Caruel, 1944) (Melitaea); Misc. ent. 41: 10
  • splendida (Mauny, 1949) (Melitaea); Lambillionea 49: 105

  • Euphydryas (Eurodryas) aurinia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38
  • Euphydryas (aurinia) aurinia; [BRU2]: 55, pl. 36, f. 19-21, 25-27
  • Euphydryas aurinia; [SPRK]; [H&R], 127; [BOW]: pl. 2, f. 6, 7; [EBW]; [BARU], #155; [MRS], 585; [Wahlberg]; [Otakar Kudrna]; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 46; Yakovlev, 2012, Nota lepid. 35 (1): 87; [FE]; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE] ; [SVF] ; [Mario Maier]
  • Eurodryas aurinia dubia; [LepIndex]; [CoL]
  • Eurodryas aurinia; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [BMAT]; [LepIndex] ; [CoL]

  500x420 (~58Kb) upperside Criptor, Devon, UK 25/05/92, Photo © Simon Coombes	550x371 (~40Kb) larva Central Europe 1990-2000, Photo © Michael Borsch	371x550 (~48Kb) pupa Central Europe 1990-2000, Photo © Michael Borsch
  595x400 (~57Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch	1024x560 (~109Kb) Russia, Moscow area, 26.05.2007, Photo © D. Smirnov	758x488 (~92Kb) larva Russia, Moscow area, 18.5.2008, Photo © D. Smirnov
  748x596 (~109Kb) upperside Russia, Moscow area (36°25'E, 56°00'N), 14.6.2009, Photo © D. Smirnov	809x652 (~98Kb) underside Russia, Moscow area, 14.6.2008, Photo © D. Smirnov	909x1133 (~275Kb) underside An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 30th June 2007 (on Senecio turczanovii), Photo © Oleg Kosterin
  1224x918 (~296Kb) underside An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 30th June 2007 (on Myosotis), Photo © Oleg Kosterin	1224x918 (~296Kb) underside An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 1st July 2007 (on Aster alpinus), Photo © Oleg Kosterin	1068x915 (~305Kb) upperside An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 1st July 2007, Photo © Oleg Kosterin
  1366x1063 (~324Kb) underside An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 1st July 2007, Photo © Oleg Kosterin	950x844 (~191Kb) upperside An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West SIberia, Russia. 3rd July 2007, Photo © Oleg Kosterin	1327x1165 (~281Kb) upperside An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West SIberia, Russia. 3rd July 2007 (on Alchemilla), Photo © Oleg Kosterin
  1004x654 (~127Kb) ♀ upperside Russia, Moscow area, 9.6.2019 (36°25'E, 56°00'N), Photo © D. Smirnov

  Larva on Plantago lanceolata, Succisa pratensis [SPRK], Knautia arvense [BARU], Scabiosa succisa, Digitalis , Plantago , Veronica , Geranium , Sambucus , Gentiana , Valeriana , Lonicera , Spiraea , Viburnum , Succisa [BRU2]: 55

  • E. a. aurinia CEU, SEU, W.Siberia

    • estonica (Petersen, 1902) (Melitaea); Beitr. Kunde Est.Liv- Kurlands 1902: 19

    • Euphydryas aurinia aurinia; [BRU2], 55; [H&R], 127; [Wahlberg]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38; [iNaturalist] ; [Heiner Ziegler]
    • Eurodryas aurinia estonica; [LepIndex]; [CoL]
  • E. a. merope (de Prunner, 1798) Alps

    • Papilio merope de Prunner, 1798; Lepid. Pedemont.: 73; TL: Valle Varaitana circa Lyaiol [Alps]
      • = Euphydryas aurinia; [FE]

    • Euphydryas (aurinia) merope; [BRU2]: 56, pl. 36, f. 28-30
    • Euphydryas merope merope; [BRU2], 56
  • E. a. provincialis (Boisduval, 1828) S.Europe, N.Africa, Asia Minor

    • aurinia provincialis (Boisduval, 1828) (Melitaea); Eur. Lepid. Index Meth.: 17; TL: Provence [France]
      • = Euphydryas aurinia; [FE]
    • ?Melitaea artemis var. provincialis Boisduval, 1840; Genera Index Eur. Lepid.: 20

    • Euphydryas (aurinia) provincialis; [BRU2]: 55, pl. 36, f. 22-24
    • Euphydryas aurinia provincialis; [H&R], 130; [iNaturalist] ; [Mario Maier]
    • Euphydryas provincialis; [MBE] ; [MBE]
    • Eurodryas aurinia provincialis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38 (?); [LepIndex]; [CoL]

    Larva on Scabiosa pratensis, S. columbaria [BRU2]: 56

  • E. a. beckeri (Lederer, 1853) Morocco (Middle Atlas, Rif Mts)

    • Melitaea beckeri Lederer, 1853; Verh. zool.-bot. Ver. Wien 2 (Abh.): 39; TL: S.Spain
      • = Euphydryas aurinia; [FE]
    • Melitaea aurinia f. iberica Oberthür, 1881; Étud. d'Ent. 6: 54; TL: Spain
    • Melitaea aurinia ellisoni Rungs, 1950; Bull. Soc. Sci. nat. Maroc 28: 145; TL: Ifrane (Morocco)
    • Melitaea aurinia kandarensis Rungs, 1950; Bull. Soc. Sci. nat. Maroc 28: 146 (= ellisoni)

    • Euphydryas aurinia beckeri; [H&R], 130; [iNaturalist]
    • Euphydryas beckeri; [LEPIFORUM]
    • Eurodryas aurinia beckeri; [BMAT]: 56, pl. 18, f. 18-25; [LepIndex]; [CoL]
    • Eurodryas aurinia ellisoni; [LepIndex]; [CoL]
    • Eurodryas desfontainii beckeri; [LepIndex]; [CoL]

    ellisoni

    600x435 (~36Kb) upperside Morocco, Imouzzèr-Kandar, Moyen Atlas, Photo © Michel Tarrier

    Larva on Lonicera implexa, L. etrusca?, L. biflora?, L. periclymenum?, (in Spain) Knautia arvensis, Succisa pratensis, Digitalis sp., Plantago sp., Veronica sp., Geranium sp. [BMAT]

  • E. a. hibernica (Birchall, 1873) Ireland

    • aurinia hibernica (Birchall, 1873) (Melitaea); Ent. Mon. Mag. 10: 154, pl. 1

    • Eurodryas aurinia hibernica; [LepIndex]; [CoL]
  • E. a. scotica Robson, 1880 Scotland

    • Melitaea[?] artemis scotica Robson, 1880; Young Naturalis 2 (58): 37
      • = Euphydryas aurinia; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 47

    • Eurodryas aurinia scotica; [LepIndex]; [LepIndex]; [CoL]
  • E. a. laeta (Christoph, 1893) C.Siberia, Altai, Sayan, Transbaikalia

    • Melitaea aurinia var. laeta Christoph, 1893; Dt. Ent. Z. Iris 6 (1): 86

    • Euphydryas (Eurodryas) laeta; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38
    • Euphydryas aurinia laeta; [BRU2], 55; [iNaturalist]
    • Euphydryas laeta laeta; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38
    • Euphydryas laeta; [iNaturalist]
    • Eurodryas aurinia laeta; [LepIndex]; [CoL]

    Larva on Succisa pratensis [BRU2]: 55

  • E. a. pellucida (Christoph, 1893) Caucasus Major

    • Melitaea aurinia var. pellucida Christoph, 1893; Dt. Ent. Z. Iris 6 (1): 86

    • Euphydryas aurinia pellucida; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38
    • Euphydryas merope pellucida; [BRU2], 56
    • Eurodryas aurinia pellucida; [LepIndex]; [CoL]
  • E. a. signifera (Kane, 1893)

    • Melitaea aurinia var. signifera Kane, 1893; Entomologist 26 (360): 158
      • = Euphydryas aurinia; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 47

    • Eurodryas aurinia signifera; [LepIndex]; [CoL]
  • E. a. praeclara (Kane, 1893) Ireland

    • Melitaea aurinia var. praeclara Kane, 1893; Entomologist 26 (360): 159

    • Eurodryas aurinia praeclara; [LepIndex]; [CoL]
  • E. a. nigra (Freke, 1903)

    • Melitaea aurinia ab. nigra Freke, 1903; Entomologist 36 (480): 110

    • Eurodryas aurinia nigra; [LepIndex]; [CoL]
  • E. a. banghaasi (Seitz, 1908)

    • Melitaea aurinia banghaasi Seitz, 1908; Gross-Schmett. Erde 1: 214, pl. 65 row c

    • Euphydryas aurinia banghaasi; [iNaturalist]
    • Euphydryas aurinia f. banghaasi;
    • Euphydryas banghaasi; Lukhtanov, 1994
    • Eurodryas aurinia banghaasi; [LepIndex]; [CoL]

    684x616 (~64Kb) ♀ upperside An alpine meadow in the valley of the left headwater of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 10th July 1998 (on Viola altaiaca), Photo © Oleg Kosterin

    527x440 (~21Kb) ♂ upperside An alpine meadow in the valley of the left headwater of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 10th July 1998, Photo © Oleg Kosterin

    762x503 (~54Kb) upperside An alpine meadow in the valley of the left headwater of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 10th July 1998, Photo © Oleg Kosterin

  • E. a. amasina (Seitz, 1908) Transcaucasia

    • Melitaea aurinia amasina Seitz, 1908; Gross-Schmett. Erde 1: 214, pl. 65 d (Stgr. i.l.)
      • = Euphydryas orientalis orientalis; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 47

    • Euphydryas aurinia amasina; [iNaturalist]
    • Euphydryas provincialis amasina; [BRU2], 56

    Larva on Cephalaria gigantea [BRU2]: 56

  • E. a. balcanica (Schawerda, 1909) Herzegowina

    • Melitaea aurinia f. balcanica Schawerda, 1909; Jber. Wien. EntVer. 19: 92

    • Eurodryas aurinia balcanica; [LepIndex]; [CoL]
  • E. a. castillana (Oberthür, 1909) Spain

    • Melitaea artemis castillana Oberthür, 1909; Étud. Lépid. Comp. 3: 226

    • Euphydryas aurinia castillana; [iNaturalist]
    • Eurodryas aurinia castillana; [LepIndex]; [CoL]
  • E. a. debilis (Oberthür, 1909) Pyrenees

    • Melitaea artemis debilis Oberthür, 1909; Étud. Lépid. Comp. 3: 228
      • = Euphydryas aurinia; [FE]
    • pyrenes-debilis [= pyrenesdebilis] (Verity, 1928) (Melitaea); Ent. Rec. 40 (6): 90

    • Euphydryas aurinia debilis; [H&R], 130; [iNaturalist]
    • Eurodryas aurinia debilis; [LepIndex]; [CoL]
  • E. a. namurcensis (Lambillion, 1909)

    • Melitaea aurinia var. namurcensis Lambillion, 1909; Rev. mens. Soc. ent. Namur. 9 (2): 21

    • Eurodryas aurinia namurcensis; [LepIndex]; [CoL]
  • E. a. alfacaria (Ribbe, 1910)

    • Melitaea aurinia v. iberica f. alfacaria Ribbe, 1910; Dt. ent. Z. Iris 23 (2): 132; TL: Sierra de Alfacar

    • Eurodryas aurinia alfacaria; [LepIndex]; [CoL]
  • E. a. morena (Ribbe, 1910)

    • Melitaea aurinia v. iberifca ab. morena Ribbe, 1910; Dt. ent. Z. Iris 23 (2): 132

    • Eurodryas aurinia morena; [LepIndex]; [CoL]
  • E. a. comacina (Turati, 1910) Italy

    • Melitaea aurinia comacina Turati, 1910; Ent. Zs. 23: 223; TL: Como, 800m

    • Eurodryas aurinia comacina; [LepIndex]; [CoL]
  • E. a. aurunca (Turati, 1910) Italy

    • Melitaea aurinia aurunca Turati, 1910; Ent. Zs. 23: 223; TL: Caserta

    • Eurodryas aurinia aurunca; [LepIndex]; [CoL]
  • E. a. volhynica (Ksienschopolski, 1911) Caucasus Major, SEEU

    • aurinia volhynica (Ksienschopolski, 1911) (Melitaea); Trudy obshch.Izsl. Volyni Zitomir 8: 35

    • Euphydryas provincialis volhynica; [BRU2], 56
    • Eurodryas aurinia volhynica; [LepIndex]; [CoL]
  • E. a. pseudomerope (Vordbrodt, 1914) Switzerland

    • Melitaea aurinia pseudomerope Vordbrodt, 1914; Die Schmett. Schweiz 2: 609

    • Eurodryas aurinia pseudomerope; [LepIndex]; [CoL]
  • E. a. anglicana (Fruhstorfer, 1917) England

    • Melitaea aurinia anglicana Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 6; TL: England, Kent
      • = Euphydryas aurinia; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 47

    • Euphydryas aurinia anglicana; [iNaturalist]
    • Eurodryas aurinia anglicana; [LepIndex]; [CoL]
  • E. a. bulgarica (Fruhstorfer, 1917) Carphathians

    • Melitaea aurinia bulgarica Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 6; TL: Bulgaria, Rila, 1700m; Rhodope

    • Euphydryas aurinia bulgarica; [BRU2], 55; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 47
    • Eurodryas aurinia bulgarica; [LepIndex]; [CoL]
  • E. a. acedia (Fruhstorfer, 1917) England

    • Melitaea aurinia acedia Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 7; TL: Wales, West England
      • = Euphydryas aurinia; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 47

    • Eurodryas aurinia acedia; [LepIndex]; [CoL]
  • E. a. volupis (Fruhstorfer, 1919)

    • Melitaea aurinia volupis Fruhstorfer, 1919; Archiv Naturg. 83 A (3): 167; TL: Tessin, Mt. Lema nr Novaggio

    • Eurodryas aurinia volupis; [LepIndex]; [CoL]
  • E. a. apiciata (Röber, 1926) Baikal

    • aurinia apiciata (Röber, 1926) (Melitaea); Ent. Mitt. 15 (3/4): 228

    • Eurodryas aurinia apiciata; [LepIndex]; [CoL]
  • E. a. rotunda (Röber, 1926) Dalmatia

    • aurinia rotunda (Röber, 1926) (Melitaea); Ent. Mitt. 15 (3/4): 228

    • Eurodryas aurinia rotunda; [LepIndex]; [CoL]
  • E. a. bicolor (Vorbrodt, 1928)

    • aurinia bicolor (Vorbrodt, 1928) (Melitaea); Dt. ent. Z. Iris 42: 20

    • Eurodryas aurinia bicolor; [LepIndex]; [CoL]
  • E. a. impunctata (Vorbrodt, 1928)

    • aurinia impunctata (Vorbrodt, 1928) (Melitaea); Dt. ent. Z. Iris 42: 20

    • Eurodryas aurinia impunctata; [LepIndex]; [CoL]
  • E. a. glaciegenita (Verity, 1928)

    • Melitaea glaciegenita Verity, 1928; Ent. Rec. 40 (3): 43
      • = Euphydryas aurinia; [FE]

    • Euphydryas glaciegenita; [Otakar Kudrna]; [MBE] ; [MBE]
    • Eurodryas aurinia glaciegenita; [LepIndex]; [CoL]
  • E. a. valentini (Nitsche, 1929)

    • Melitaea aurinia var. valentini Nitsche, 1929; Verh. zool.-bot. Ges. Wien 78 (S.B): 14
      • = Euphydryas aurinia; [FE]

    • Eurodryas aurinia valentini; [LepIndex]; [CoL]
  • E. a. kricheldorfi (Collier, 1933) Spain

    • aurinia kricheldorfi (Collier, 1933) (Melitaea); Ent. Rundsch. 50: 54

    • Eurodryas aurinia kricheldorfi; [LepIndex]; [CoL]
  • E. a. infraflava Verity, 1950 Italy

    • aurinia infraflava Verity, 1950; Le Farfalle diurn. d'Italia 4: 71

    • Eurodryas aurinia infraflava; [LepIndex]; [CoL]
  • E. a. infraochrea Verity, 1950 Italy

    • aurinia infraochrea Verity, 1950; Le Farfalle diurn. d'Italia 4: 71

    • Eurodryas aurinia infraochrea; [LepIndex]; [CoL]
  • E. a. sesquiargentea Verity, 1950 Italy

    • aurinia sesquiargentea Verity, 1950; Le Farfalle diurn. d'Italia 4: 72

    • Eurodryas aurinia sesquiargentea; [LepIndex]; [CoL]
  • E. a. basiconfluens Lempke, 1955 Holland

    • Euphydryas aurinia f. basiconfluens Lempke, 1955; Tijdschr. Ent. 98: 337; TL: Wageningen

    • Eurodryas aurinia basiconfluens; [LepIndex]; [CoL]
  • E. a. inframaculata Lempke, 1955 Holland

    • Euphydryas aurinia f. inframaculata Lempke, 1955; Tijdschr. Ent. 98: 337; TL: Apeldoorn

    • Eurodryas aurinia inframaculata; [LepIndex]; [CoL]
  • E. a. nigromaculata Lempke, 1955 Holland

    • Euphydryas aurinia f. nigromaculata Lempke, 1955; Tijdschr. Ent. 98: 337; TL: van Wageningen

    • Eurodryas aurinia nigromaculata; [LepIndex]; [CoL]
  • E. a. barraguei (Betz, 1956) Algeria

    • Melitaea aurinia barraguei Betz, 1956; Revue fr. Lép. 15: 144; TL: Berrouaghia (Algeria)

    • Eurodryas aurinia barraguei; [BMAT]: 56, pl. 18, f. 26-29; [CoL]
  • E. a. celina Krzywicki, 1967 Poland

    • Euphydryas aurinia celina Krzywicki, 1967; Ann. zool. Warsz. 25 (1): 111

    • Eurodryas aurinia celina; [LepIndex]; [CoL]
  • E. a. lucana Hartig, 1968 Italy

    • aurinia lucana Hartig, 1968; Reichenbachia 12 (1): 1

    • Eurodryas aurinia lucana; [LepIndex]; [CoL]
  • E. a. jungei Gross & Ebert, 1975 N.Iran

    • Euphydryas aurinia jungei Gross & Ebert, 1975; J. ent. Soc. Iran, suppl. 1: 34, f. 40-41; TL: Elburs
  • E. a. tiragalloi Balletano & Toso, 1976 Italy

    • aurinia tiragalloi Balletano & Toso, 1976; Doriana 5 (225): 2

    • Eurodryas aurinia tiragalloi; [LepIndex]; [CoL]
  • E. a. altivolans Tuzov, 2000 Altai, Sayan, S.Siberia

    • Euphydryas merope altivolans Tuzov, 2000; ; TL: Altai, Severochujskij Mtns, Sunko Mt

    • Euphydryas aurinia altivolans; Yakovlev, 2012, Nota lepid. 35 (1): 87
    • Euphydryas merope altivolans; [BRU2], 56
  • E. a. avis (Lastuchin, 2008)

    • aurinia avis (Lastuchin, 2008) (Eurodryas)

    • Euphydryas aurinia avis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38
• Euphydryas orientalis (Herrich-Schäffer, [1851]) -. See [maps]

  • Melitaea orientalis Herrich-Schäffer, [1851]; Syst. Bearb. Schmett. Europ. 1 (10): (ii) pl. 56, f. 265-266; TL: [Amasya, Istanbul, Turkey]

  • Euphydryas (aurinia) orientalis; [BRU2]: 56, pl. 36, f. 31-36
  • Euphydryas orientalis; [Wahlberg]; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 47; [iNaturalist] ; [LEPIFORUM]
  • Eurodryas orientalis; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [LepIndex] ; [CoL]
  • E. o. orientalis Turkey, Transcaucasia

    • Euphydryas orientalis orientalis; [BRU2], 56
  • E. o. sareptensis (Staudinger, 1878) S.Ural

    • Melitaea artemis v. sareptana Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 8 (nom. nud.?)
    • Melitaea aurinia v. sareptana Staudinger, 1871; Cat. Lepid. eur. Faunegeb. (Edn. 2): 17 (nom. nud.?)
      • = Melitaea aurinia v. orientalis; Staudinger, 1871, Cat. Lepid. eur. Faunegeb. (Edn. 2): 17
    • Melitaea aurinia var. sareptensis Staudinger, 1878; Horae Soc. Ent. Ross. 14: 265; TL: Sarepta

    • Euphydryas aurinia sareptana; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38
    • Euphydryas aurinia sareptensis; [iNaturalist]
    • Euphydryas orientalis sareptensis; [BRU2], 56
    • Euphydryas sareptana; [BOW]: pl. 194, f. 20
    • Eurodryas aurinia sareptana; [LepIndex]; [CoL]
  • E. o. emba (Fruhstorfer, 1917) N.Kazakhstan

    • Melitaea aurinia emba Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 7; TL: Emba, Uralsk
      • = Euphydryas aurinia sareptana; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38

    • Euphydryas orientalis emba; [BRU2], 56
• Euphydryas asiatica (Staudinger, 1881) Turkestan. See [maps]

  • Melitaea aurinia var. asiatica Staudinger, 1881; Stettin ent. Ztg 42 (7-9): 287; TL: "Ala Tau" [Dzhungarsky Alatau Mts., Kazakhstan]
  • Melitaea aurinia var. asiatica Alphéraky, 1882; Horae Soc. ent. Ross. 16 (3-4): 400 (Stgr. in litt.)

  • Euphydryas (Eurodryas) asiatica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38
  • Euphydryas (aurinia) asiatica; [BRU2]: 57, pl. 37, f. 1-6
  • Euphydryas asiatica; [Oleg Kosterin]; [iNaturalist] ; [LEPIFORUM]
  • Euphydryas aurinia asiatica; [Wahlberg]; Yakovlev, 2012, Nota lepid. 35 (1): 87; [iNaturalist]

  533x400 (~60Kb) ♂ upperside A meadow in the lower valley of the Kora River about 10 km upstream of the Tekeli City (1300 m above sea level), the North Dzhungarskiy Alatau mountain chain (NE part of Tien Shan), Kopal District, Taldy-Kurgan Province, SE Kazakhstan. 13th June 1993, Photo © Oleg Kosterin

  • E. a. asiatica Tarbagatai, Saur, Dzhungarsky Alatau

    • Euphydryas asiatica asiatica; [BRU2], 57; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38
  • E. a. alexandrina (Staudinger, 1887) Tian-Shan

    • Melitaea aurinia var. alexandrina Staudinger, 1887; Stettin Ent. Ztg 48 (1-3): 52

    • Euphydryas alexandrina; [Wahlberg]
    • Euphydryas asiatica alexandrina; [BRU2], 57; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38; [iNaturalist]
    • Eurodryas alexandrina; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [LepIndex] ; [CoL]
  • E. a. narina (Oberthür, 1909) Inner Tian-Shan

    • Melitaea artemis var. narina Oberthür, 1909; Étud. Lépid. Comp. 3: 230

    • Euphydryas asiatica narina; [BRU2], 57; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38
• Euphydryas sibirica (Staudinger, 1871) -. See [maps]

  • Melitae artemis v. sibirica Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 7

  • Euphydryas (Eurodryas) sibirica; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38
  • Euphydryas (aurinia) sibirica; [BRU2]: 57, pl. 37, f. 7-15
  • Euphydryas sibirica; [iNaturalist] ; [LEPIFORUM]
  • Melitaea aurinia v. sibirica; Staudinger, 1871, Cat. Lepid. eur. Faunegeb. (Edn. 2): 17

  1072x918 (~257Kb) ♀ upperside A steppe on the Kurota River left bank at its mouth, 12 km SW of Ongudai village, Ongudai District, Altai Republic, West Siberia, Russia. 7th July 2005, Photo © Oleg Kosterin

  1177x918 (~269Kb) ♀ upperside A steppe on the Kurota River left bank at its mouth, 12 km SW of Ongudai village, Ongudai District, Altai Republic, West Siberia, Russia. 7th July 2005, Photo © Oleg Kosterin

  1196x909 (~309Kb) ♀ upperside A steppe on the Kurota River left bank at its mouth, 12 km SW of Ongudai village, Ongudai District, Altai Republic, West Siberia, Russia. 7th July 2005, Photo © Oleg Kosterin

  Larva on Veronica [RFEL]

  • E. s. sibirica Transbaikalia

    • Euphydryas aurinia sibirica; [Oleg Kosterin]
    • Euphydryas sibirica sibirica; [BRU2], 57; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38; [iNaturalist]
  • E. s. eothena (Röber, 1926) Amur, Ussuri

    • sibirica eothena (Röber, 1926) (Melitaea); Ent. Mitt. 15 (3/4): 228
    • Euphydryas tjutjujensis Higgins, 1950; Trans. R. ent. Soc. Lond. 101 (12): 466

    • Euphydrya davidi tjutjujensis; [BARU](note) ,
    • Euphydryas sibirica eothena; [BRU2], 57
    • Euphydryas sibirica tjutjujensis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38

    Larva on Scabiosa lachnophylla [BRU2]: 57

  • E. s. davidi (Oberthür, 1881) N.China, Tuva?, Mongolia?

    • Melitaea artemis var. davidi Oberthür, 1881; Étud. d'Ent. 6: 52 (footnote); TL: N.China, Peking
    • Melitaea aurinia var. mandschurica Staudinger, 1892; Dt. Ent. Z. Iris 5 (2): 323 (preocc. Melitaea athalia var. mandschurica Fixsen, 1887)
      • = Euphydryas aurinia davidi; [Wahlberg]
    • mandschura (Bryk, 1946) (Melitaea); Ark. Zool. 38 A (3): 47 (repl. Meliaea aurinia var. mandschurica Staudinger, 1892)

    • Euphydryas aurinia davidi; [Wahlberg]
    • Euphydryas aurinia mandschurica; [MRS], 585
    • Euphydryas davidi; [BARU], #156
    • Euphydryas sibirica davidi; [BRU2], 57; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38 (?); [iNaturalist]

    800x600 (~151Kb) upperside Wet ground at the pool in the pine forest called Tsasucheiskii Bor at the village Nizhnii Tsasuchei, Onon District, SE Chita Province, Transbaikalia, Siberia, Russia. 22nd June 1995, Photo © Oleg Kosterin

    534x800 (~117Kb) ♂ ♀ underside A meadow on the bank of Lake Butyvken, surrounded by a magnificent pine forest within a steppe zone called Tsasucheiskii Bor, 10 km SW of the village Nizhnii Tsasuchei, Onon District, SE Chita Province, Transbaikalia, Siberia, Russia. 25th June 1995 (on Oxytropis myriophylla), Photo © Oleg Kosterin

    450x600 (~56Kb) underside A meadow on the bank of Lake Butyvken, surrounded by a magnificent pine forest within a steppe zone called Tsasucheiskii Bor, 10 km SW of the village Nizhnii Tsasuchei, Onon District, SE Chita Province, Transbaikalia, Siberia, Russia. 25th June 1995 (a congregation), Photo © Oleg Kosterin

  • E. s. tenebricosa (Bang-Haas, 1927) Gansu

    • sibirica tenebricosa (Bang-Haas, 1927) (Melitaea); Horae Macrolep. Palaearct. 1: 51

    • Euphydryas sibirica tenebricosa; [BRU2], 57
  • E. s. phyllis Hemming, 1941 N.Korea

    • sibirica phyllis Hemming, 1941; Proc. R. ent. Soc. Lond. (B) 10 (10): 207 (repl. koreana Collier, 1933)
    • koreana (Collier, 1933) (Melitaea); Ent. Rundsch. 50: 54 (preocc.)
    • discalis (Bryk, 1946) (Melitaea); Ark. Zool. 38 A (3): 47

    • Euphydryas aurinia discalis; [iNaturalist]
    • Euphydryas sibirica phyllis; [BRU2], 57
• Euphydryas desfontainii (Godart, 1819) NAF, Spain, Pyrenees. See [maps] Dientes gualdos Spanish Fritillary

  • Argynnis desfontainii Godart, 1819; Encyclopédie Méthodique 9 (1): 278; TL: N.Africa, Spain
  • Melitaea artemis var. desfontainesii Boisduval, 1840; Genera Index Eur. Lepid.: 20
  • zapateri Higgins, 1950; Trans. R. ent. Soc. Lond. 101 (12): 482

  • Euphydryas desfontainii boumalnei; [iNaturalist]
  • Euphydryas desfontainii iberae; [iNaturalist]
  • Euphydryas desfontainii; [H&R], 131; [BOW]: pl. 2, f. 9; [Wahlberg]; [FE]; [Otakar Kudrna]; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE] ; [Mario Maier]
  • Eurodryas desfontainii; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [LepIndex] ; [CoL]

  Larva on Knautia [H&R]

  • E. d. desfontainii Morocco (Rif Mts, Middle Atlas), W.Algeria

    • Melitaea desfontainii var. gibrati Oberthür, 1922; Étud. Lépid. Comp. 19 (1): 46, (2): pl. 532, f. 4418-4422; TL: Morocco

    • Euphydryas desfontainii desfontainii; [H&R], 131
    • Eurodryas desfontainii desfontainii; [BMAT]: 56, pl. 18, f. 30-37
    • Eurodryas desfontainii gibrati; [LepIndex]; [CoL]

    gibrati

    600x435 (~77Kb) ♂ ♀ Morocco, Tizi-n-Tretten, Moyen Atlas, Photo © Jean Delacre

    600x435 (~61Kb) ♂ ♀ underside Morocco, Tizi-n-Tretten, Moyen Atlas, Photo © Jean Delacre

    600x435 (~58Kb) ♂ ♀ upperside Morocco, Ito, Moyen Atlas, Photo © Michel Tarrier

    Larva on Knautia arvensis [BMAT]

  • E. d. baetica (Rambur, 1858) Spain

    • desfontainii baetica (Rambur, 1858) (Melitaea); Cat. syst. Lépid. Andalousie (1): 11, pl. 1, f. 3-4
      • = Euphydryas desfontainii; [FE]

    • Euphydryas aurinia baetica; [iNaturalist]
    • Euphydryas desfontainii baetica; [H&R], 131
    • Eurodryas desfontainii baetica; [LepIndex]; [CoL]
  • E. d. ibarrae Lajonquiére, 1965 Spain

    • desfontainii ibarrae Lajonquiére, 1965; Bull. mens. Soc. linn. Lyon 34 (2): 47

    • Eurodryas desfontainii ibarrae; [LepIndex]; [CoL]
  • E. d. quevedoi (Agenjo, [1970]) Spain

    • desfontainii quevedoi (Agenjo, [1970]) (Melitaea); Graellsia 25: 169

    • Eurodryas desfontainii quevedoi; [LepIndex]; [CoL]
  • E. d. moralesi Gómez Bustillo, 1973 Spain

    • desfontainii moralesi Gómez Bustillo, 1973; SHILAP Revta Lep. 1 (1-2): 26

    • Eurodryas desfontainii moralesi; [LepIndex]; [CoL]
  • E. d. martae

    • Euphydryas desfontainii martae; [iNaturalist]

    800x580 (~68Kb) upperside Morocco, Djebel Azourki, Haut Atlas, Photo © Jean Delacre